A Checklist and Generic Key to the Tabanidae (Diptera) of Perti with Special Reference to the Tambopata Reserved Zone, Madre de Dios q

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1 23: Rev. per. Ent. 27: Diciembre 1984 (30 Setiembre 1985) A Checklist and Generic Key to the Tabanidae (Diptera) of Perti with Special Reference to the Tambopata Reserved Zone, Madre de Dios q Richard C. Wilkerson3 and G. B. FairchId SUMMARY Two hundred and twenty-eight species of horse flies (Diptera: Tabanidae) are documented for Peru, with 73 species collected from a single lowland forest site in southeastern Peru. No other site in the Neotropics has as many reported species. A key to the subfamilies, tribes, and genera of the known Peruvian species is also included. SUMARIO Se cita 228 especies de tibanos registradas para el Peru, con 73 especies colectadas en una localidad de bosque amazonico bajo de1 sudeste de1 pais. Ningun otro lugar de1 Neotropico presenta tantas especies reportadas hasta el momento. Se in&ye asimismo una clave para las subfamilias, tribus y generos de tabanos conocidos para el Peru. INTRODUCTION Compilation of this list of Peruvian Tabanidae began initially with a survey of the Tambopata Reserved Zone. To the list of the Tabanidae of this Reserve we have added all other records known to us from Peru. The Reserve is an area of approximately 5,500 ha located about 39 km (air) from Puerto Maldonado, Madre de Dios (12 15 S lat. and W long). It is in a Subtropical Moist Forest life zone (ONERN 1976) with an annual rainfall of mm and mean annual biotemperature of C. The average altitude of the reserve is 300 m, and the topography is generally flat with some small hills and oxbow lakes. Except for the immediate environs of the tourist hotel and scientific base (the Explorer s Inn ) the area is made up of virgin forest. There are several distinct forest types that are dictated by the type and drainage of the soil substrate. They range from flood plain and bamboo forest to well drained forests with tall canopies (40 m or more). The Tabanidae (horse flies, deerflies, tabanos ) are a diverse family of true flies with worldwide distribution. There are approximately 3500 described species, of which 1100 are named from the Neotropics. 1 Recibido para publicacicq 31 mayo Florida Agricultural Experiment Station Journal Series No Florida Department of Agriculture, Division of Plant Industry, P. 0. Box 1269, Gainesville, Florida 32602, U.S.A. 37 PERUVIAN HORSE FLIES Previous studies on the Tabanidae of Peru have consisted principally of scattered descriptions of new species or superficial fauna1 lists derived from the literature. Soukup (1945) gives a brief list of the Diptera of Peru, the Tabanidae largely culled from the literature up to 1940, especially the Catalogue of Kroeber (1934). He gives very few specific iocalities, though listing 164 names. Kroeber (1951), in a report on an expedition to south Peru from the Hamburg Museum, gave a list of 20 genera and

2 38 REVISTA PERUANA DE ENTOMOLOGIA. Vol species known to him from Peru, often with localities and synonymy. To this list he added 14 species, with localities, taken by the expedition. Philip (1958, 1960) described a number of new species from Peru, and discussed others, mostly from Tingo Maria in the Department of Huanuco. Finally, Carrasco (1972) published a list of 163 species for Peru of which 29 species from southern Peru were collected by himself and determined by Fairchild; ten of these were not determined to species. Unfortunately, the localities copied from the literature, and those on specimens he or Fairchild determined are not distinguished. Other than suggesting that the Peruvian fauna is a rich one, these previous lists of names are often of dubious validity and uncertain ascription. SOURCES OF SPECIMENS We have been fortunate to study several extensive collections of Tabanidae from Peru. Fairchild (G.B.F.) h as seen much of the material reported by Philip (C.B.P.) collected by Ross and Michelbacher in Tingo Maria and now in the California Academy of Sciences (CAS), as well as material in the Museum of Comparative Zoology in Cambridge, Mass. (MCZ) collected by WK. Weyrauch and the specimens in L. L. Pechuman s (LLP) collection in Cornell, New York, collected in Avispas, Madre de Dios and Quincemil, Cuzco, by Luis Petia. Fairchild also made a small collection in the Rimac Valley above Lima in By far the largest collections seen from Peru are those from the Rio Tambopata Reserve near Puerto Maldonado, made in 1979 by J.B. Heppner and by R. C. Wilkerson (RCW) twice in Other than these collections, we have depended mainly on specimens in Fairchild s collection or records derived from determinations made by him from various sources. He also studied the types of most neotropical species, including those from Peru, reported in a series of publications (1953, 1961, 1966a, b, c, d, 1967) culminating in his Catalogue (1971). ~"" ~Y*aL*v*IJ ALL.-_,..;~+:~.w an&/or lucaliui1~ I,.:_-- of C&Cl sources of specimens seen by Fairchild included in this list follow: American Museum of Natural History (AMNH), New York; Florida State Collection of Arthropods (FSCA), Gainesville, Florida; Zoologisches Museum der Math. - Naturwissenschaftlichen Facultat der Humboldt Universitat zu Berlin, Berlin, East Germany; San Diego State University, San Diego CA, USA; British Museum (Natural History) (BMNH), London; National Museum of Natural History (USNM), Washington, D.C.; Canadian National Collection (CNC), Ottawa; Ohio State University, Columbus, Ohio; Museum National d Histoire Naturelle, Paris, France; Staatliches Museum fur Tierkunde, Dresden, East Germany; Zoologische Sammlung des Bayerischen Staates, Munich, West Germany; Museo de Historia Natural, Universidad National Mayor de San Marcos, Lima (MUSM); Rijk smuseum van Natuurlijke Historie, Leiden, Netherlands. GENERAL ECOLOGY Most horse flv females require vertebrate blood for completion of at least the second and subsequent gonotrophic cycles. Males do not feed on blood but on nectar. Females also utilize sugars as an energy source. Horse flies have a large range of hosts including mammals, birds and large reptiles. Horse fly larvae are usually predacious and are found mostly in aquatic or semi-aquatic habitats. For further general information on the family Tabanidae in the neotropics see the following: Barrett0 (1946), morphology; Fairchild (1969), generic classification; Fairchild ( 197 l), catalogue; Fairchild (198 1)) general information with annotated references; Krinsky (1976), disease agents transmitted by tabanids; Goodwin and Murdoch (1974), immatures; and Wilkerson et al. (1985), mating behavior. For purposes of describing horse fly distribution, we consider Peru to consist of three main regions: 1. The Pacific coast, and valleys of rivers flowing into the Pacific. This area is mainly desert with intensive irrigated cultivation along the short coastal rivers. 2. The highlands, extending from 1500 m to over 4000 m, densely populated and intensely cultivated since ancient times, either irrigated or dependent on low to moderate rainfall, but with little original woody vegetation remaining. 3. The eastern lowlands and foothills. A region, mostiy beiow i5ov m, sparseiy popuiated, wrth high rainfall and where not cleared for agriculture, extensively covered by high tropical to subtropical forest. The first region has been little collected, and most of the records of Tabanidae have come from the vicinity of Lima or southward to Tacna. We have encountered almost no records of Tabanidae from the northern half of the country at any elevation. The few species reported have been either precinctive, stragglers from adjacent highlands, or widespread species.

3 Dloembre 1984 WILKERSON & FAIRCHILD: TABANIDAE OF TAMBOPATA 39 The fauna of the Peruvian highlands is distinct, consisting largely of species of Scaptia, Scione and Dasybasis, the first and last showing close relationships to Chilean forms, while &zone seems to have its main center in Ecuador. Most collecting has been from the southern part of the country, above Lima and Arequipa and around Lake Titicaca. Unlike the other regions of Peru, relatively extensive collections of Tabanidae have been made at a few localities in the eastern lowlands, primarily at: Tingo Maria, Department of Huanuco; Quincemil, Department of Cuzco; and Tambopata and Avispas, Department of Madre de Dios. Smaller collections are available from Iquitos, Department of Loreto and Satipo, Department of Junin. These collections reveal the predominance of an essentially Amazonian fauna. Species found in Western Ecuador and Colombia north into Panama are generally absent, while many species from Tambopata extend north into the Guianas and east to the mouth of the Amazon. CONTENT AND ARRANGEMENT OF LIST The following list includes only species that have: been seen by us; for which we have firsthand published information by credible witnesses; or which have a Peruvian type locality. The arrangement of genera and species follows Fairchild s catalogue C 197 1). Specimens from the reserve may be found in the collections of the authors (at FSCA), in the Museo de Historia Natural (Lima) or in the United States National Museum of NaturaI History. The source(s) of a record are given in the list. They may include a literature reference, and/or the presumed depository(ies) of the specimens examined by Fairchild. A record followed only by Fairchild s initials (GBF) indicates the specimen(s) are in his collection (now at FSCA). Stypommisa (Styphocera) antennina Philip 1969 is not keyed, as we are unsure of its position, We have studied the type from Peru and a pair from Ecuador. The species will probably key out to Dasybasis (Agelanius) in the key. It somewhat resembles D. (A.) Zautus Hine, though smaller and with very hairy antennae. The Ecuadorian specimens have a normal 4- annulate antenna1 style, so we doubt that the 3- annulate style of the holotype is of generic significance. We see no resemblance to any Stypommisa. KEY TO SUBFAMILIES, TRIBES AND GENERA OF PERUVIAN TABANIDAE (See Figs. A-J) Hind tibiae with paired terminal spurs or spines, rarely absent or difficult to see 2 Hind tibiae without such paired spines Tabaninae 10 Ninth tergite undivided, a large sclerotized shield in males, a transverse bar in females. Sclerotized section of spermathecal ducts of female without a cup-like expansion. Vein SC bare above and below. Antenna1 flagellum of 7 or 8 distinct flagellomeres... Pangoniinae 3 Ninth tergite divided in males, usually so in females. Sclerotized section of spermathecal ducts without a cup-like expansion. Basal 3 or 4 flagellomeres fused into a basal plate... Chrysopsinae 9 Style of male genitalia bifid. Eyes bare. First posterior cell (5th R) nearly always closed. A short to long appendix generally present at fork of third vein CR,+,)......_... Pangoniini 4 Style of male genitalia simple, pointed. Eyes nearly always pilose. First posterior cell open or closed, rarely with an appendix.._ Scionini 5 Proboscis with theta and labella shiny sclerotized, the latter slender and with the pseudotracheae reduced or Iacking. Wings hyaline or weakly tinted, unicolorous......, Esenbeckia (Probosmides). Proboscis with theta pollinose, the labella broad and shiny, pseudotracheae present. Wings often strongly patterned, bicolored or tricolored... Esenbeckia (Esenbeckia). Antennae with a strong dorsal pointing hook on first flagellar segment, and shorter dorsal protuberances on next five flagellar segments. First posterior cell closed far from wing margin by an anterior bending of vein Mi. Frontoclypeus inflated and shiny... Pityocera (Elaphella) Antennae simple, rarely with short dorsal and ventral extensions on flagellar segments. First posterior cell closed by a posterior bending of vein R5, or open. Frontoclypeus various Both first (5th R) and fourth (3rd M) wing cells closed, the wings often spotted or patterned. Frontoclypeus pollinose Scione Fourth posterior cell always open, first open or closed. Frontoclypeus often partly bare... 7 Proboscis short, rarely exceeding head height, the labella large and largely membranous. Palpi long, over l/2 length of proboscis. Frontoclypeus short, not con-

4 40 REVISTA PERUANA DE ENTOMOLOGIA Vol. 27 icdy produced and always pollinose..._......i... Scap tia (Scap tia) Proboscis long to very long, at least twice head height and often longer than body, the labella slender and sclerotized. Palpi slender, never over l/4 length of pr&- oscis. Frontoclypeus often much produced, conical, generally at least partly bare and shiny Fidena 8 8. Legs with short pilose tibiae, the femora short or long pilose Fidena (Laphriomyia) 9. Wing with first posterior cell closed and petiolate and wing entirely black except for a clear streak in discal cell and commashaped mark in axillary cell. Eyes pilose. Whole insect almost entirely black and shiny, closely mimicking a wasp Rhynomyzini Betrequia Wing with first posterior cell wide open and never so extensively black. Eyes bare, not mimicking Hymenoptera.....,... Chrysopsini Chvsops 10. Basicosta generally bare, lacking strong setae. Or if setae are present these sparser than on adjoining Costa, and often with one or more of the following characters: vestiges of ocelli or an ocellar tubercle, sclerotized labella of proboscis, strongly patterned wings, inflated tibiae, a dorsal spine on basal plate of antenna; see couplet Diachlorini 11 Basicosta with strong setae as dense as those on costa and very rarely with any of the above characters... Tabanini With one or both of the following character: basal plate of third antenna1 segment with an acute dorsal spine or tooth; labella of proboscis compact and with at least some shiny sclerotized areas Without either of the above characters, the basal plate at most obtusely angled above and the labella wholly pollinose Basal plate of third antenna1 segment with at most an obtuse angle, 90 or more, never excessively long nor with first segment cylindrical and elongate. Labella of proboscis pa+ or who& c&mt;7j 1_*_1 vc-_u Basal plate with a long or short spine or acute angle, or greatly elongate, both style and first segment unusually long. Labella scerotized or not First antenna1 segment shiny and globose, moderately to markedly inflated. Subcallus inflated and shiny First antenna1 segment normal, subcallus pollinose Vein & of wing bent abruptly forward so that cell 3rd R is somewhat narrowed at wing marain. All tibiae usuallv more or less inflated. Wings and body largely black or yellow, the wing apex always hyaline. Third antenna1 segment slender, the plate with small angle close to base dorsally. Palpi slender, pollinose. Eyes with lower half irregularly speckled, upper half unicolorous..._. Bolbodimyia Vein & not bent forward. Tibiae slender. Palpi inflated and shiny. First antenna1 segment very greatly inflated, black; third segment elongate, the style 3 times as long as basal plate, the latter with strong dorsal tooth. Frontoclypeus and genae extensively bare and shiny. Wing with anterior half black. Wasp-like species with basal callus as wide as frons... Querbetia Frontal calms absent. Basal plate broad with obtuse dorsal angle. Labella wholly scerotized. Pale yellowish or greenish unicolorous stout flies with wings hyaline or with small discrete black spots......*...*... C~blorotabanus Frontal callus present. Larger species with unstriped thorax, not mimetic. Frontal cal-!us small and rounded, usually with a slender upper ridge-like extension. Subcallus bare or pollinose. Eyes unicolorous, green or bronzy in life. Palpi more slender, pointed, over half length of proboscis. Labella largely or wholly sclerotized. Wings generally with a dark pattern, sometimes hyaline. Abdomen of female normal Phaeotabanus Frontal callus as wide as frons, or if not, frons widened below, callus rounded and protuberant and bare areas on face. Eyes usually with at least a median dark transverse stripe, generally with several green or purple stripes, rarely unicolorous. Slender species, the tibiae slender, abdomen never green...f Frontal callus narrower than frons; if not, then triangular and extended upward in a ridge, the lower angles barely touching eyes, or tibiae incrassate or abdomen greenish. Eyes unicolorous or bicolored, very rarely striped..._ A *.*LL*L**~~ n+pnnnn Xkyf!oilg, *iist c 2 StZgiZ2iitS fic3t- ably elongate, the first segment subequal to basal plate of third segment. Style generally hirsute and much exceeding basal plate, stout. Dorsal spine of basal plate moderate to absent. Frontoclypeus and genae largely or wholly bare and shiny, at least the first much inflated. Abdomen generally constricted basally, wasp-like. Wings with fore border broadly infuscated, rarely with an additional dark diagonal fascia. Labella of proboscis partly sclerotized Acanthocera (Acantbocera)

5 . Diciembre 7984 WILKERSON & FAIRCHILD: TABANIDAE OF TAMBOPATA 41 Figs. A-J. A, head, Fidena auribarba. B, head, Chrysups varlegagala. C, head, side view, DasycbeIa badza. D, apex of hind tibia of Fidena flaviperznis showing bare basicosta and setose base of costal vein. F, base of wing of Tabanus nebulosus showing setose basicosta and base of costal vein. G, wing of Scitine costaniana with the cells labeled according to Curran. Note the closed and petiolate first and fourth posterior cells and anal cell. H, wing of Tabanus pseudonrlus with the veins named according to Comstock-Needham. Note the appendix of fork of third vein and coarctate first posterior cell. I side view of thorax of PoeciIoderas quadrc)unctatus. J, dorsal view of thorax and first 2 abdominal segments of Tabanus iebulosus.

6 42 REVISTA PERUANA DE ENTOMOLOGIA Vol Antennae not unusually long, the first segment notably shorter than basal plate. Dorsal spine short to very long. Style not unusually hirsute, rarely longer than plate, slender. Abdomen slightly if at all constricted. Wings various, rarely unpatterned. Labella often entirely sclerotized Dicbelacera s Labella of proboscis wholly scerotized and shiny. Wings with a diagonal dark fascia from apex to fifth posterior cell. Eyes with one or more transverse bands. Scutellum nearly always darker than adjoining mesonotum; the latter usually transversely banded.... Dichelacera (Dicbelacera) Labella partly pollinose. Eyes unicolorous. Center of frontoclypeus and subcallus bare and shiny. Wings yellowish tinted, with area beyond fork of third vein and apex of discal cell faintly dusky. Mesonotum striped, the scutellum black Dicbelacera (Idiocbelacera). Tibiae all slender. Tubercle at vertex generally distinct and prominent, rarely obsolete. Palpi slender to thread-like. Proboscis generally with small compact labelia, wholly or partly scerotized, rarely wholly membranous. Antennae slender, the dorsal spine slender, pointed, rarely shorter than basal plate. Frons usually narrow, the callus clavate or ridge-like, rarely otherwise. Wings nearly always patterned, rarely lightly tinted, never wholly clear Catachlorops 20 Tibiae inflated, or tubercle at vertex absent, or palpi inflated, or dorsal antenna1 spine clubbed or very short, or frons broad First posterior cell closed or strongly coarctate. Frons very narrow, the callus ridge-like. Wings with dark pattern which leaves area around apices of basal cells or most of discal cell, and apex or oval spots in 1st R and 3rd R clear or paler. Antennae with long slender tooth, its apex often recurved. Labella wholly sclerotized. Palpi very slender... Catacblorops (Psalidia) Fit-cc A ll I nnctf3-int r if sn_mpwfiat coarctate, then wings otherwise Wings with a dark discal patch which includes all of discal cell, the latter sometimes fenestrate, or wings wholly black with hyaline apex, the basal cells hyaline or partly or wholly dark. Frontal callus often clavate. Labella often partly pollinose. Thorax at most faintly striped Catacblorops (Catacblorops) Wings not as above Stout species with relatively broad frons, whollv pollinose labella. moderatelv stout &l nrwn * -I palpi, and chunky antennae Catacblorops (Hadrocblorops) Slender species with narrower frons, partly sclerotized labella, slender palpi and slender antennae Catacblorops (Ampbicblorops) Palpi basally inflated. Proboscis short, heavy, labella wholly sclerotized. Stout species with proportionately short stout antennae _ Palpi long, slender, generally exceeding antennae. Proboscis long to very long, slender, the labeila pollinose. Not stout chunky species, the antennae slender. Brown species with a dark wing pattern which leaves area around ends of basal cells and apex of wing hyaline or paler 25 Dorsal tooth on third antenna1 segment short, an acute angle to a short spine, seldom exceeding end of basal plate. Fore tibiae slender to moderately inflated, remaining tibiae slender. Wings glass-clear to slightly smoky. Abdomen and appendages often greenish in life. Not bee-like species... C7yptotyhs Dorsal tooth always exceeding end of basal plate, sometimes nearly reaching apex of style, often clubbed. Fore tibiae always incrassate, remaining tibiae generally also inflated. Wings various, never wholly hyaline or uniformly tinted, generally with black or contrasting pattern, often resembling bees... Stibasoma (Stibasoma) Basal plate of antenna with one or two long dorsal spines, one or both of which reach end of basal plate, their opposing surfaces fringed with long erect hairs. Style very short, hardly more than one-third length of basal plate. Proboscis exceeding head height, the labella small, pollinose. Eyes bare Dasycbela (Triceratomyia) Basal plate with but one dorsal spine, without fringing hairs. Antennae generally shorter, the style proportionately longer. Proboscis as above or shorter. Eyes often pilose Dasycbela (Dasycbela) nisral -1 _UI r.-11 A1 L nf vrinn....ab n~rtmmd -1-*-Y.. -- ho J antm-inr --*--- bending of vein M,. Small blackish species with wings largely black to beyond discal cell, inflated shiny palpi, inflated tibiae, and elongate first antenna1 segment Lepiselaga Discal cell normal Wing basally black or heavily tinted to ends of basal cells, remainder hyaline. Whole face and entire body including legs, black and shiny. Palpi strongly inflated. Third antenna1 segment subcylindrical, without marked dorsal angle...

7 Diciembre 1984 WILKERSON & FAIRCHILD: TABANIDAE OF TAMBOPATA Himantostyhs Wing otherwise Notopleural lobes bluntly conical, protuberant. Fore tibiae inflated, hind tibiae flattened and with a fringe of long hairs. Subcallus bare. Frontal callus broadly clavate. Wings with a narrow abbreviated discal band below stigma. Eyes unicolorous Eutabanus Notopleural lobes normal, tibiae usually slender, the hind pair never flattened. 29 Mesopleuron and mesosternum darker than adjoining sclerites, the mesopleuron shiny pearly pollinose. Wings usually with dark marking, the apex often blackish. Eyes with characteristic pattern of green spots and stripes, resembling Chrysops......_...._..._.... Diachlorus Mesopleuron and mesosternum concolorous with adjoining sclerites, not pearly pollinose. Wings various. Eyes unicolorous or banded, but not as above Eyes bare with at least two transverse bands in life, light on dark purplish or blackish. Mostly small species with bare eyes, moderately broad frons with often a median dark-haired patch, and rounded or square callus generally as wide as frons. (If frons very narrow, callus ridge-like and eyes unbanded, see Stypommisa couplet 40)... Stenotabanus s Eyes with at most a single dark median stripe, generally unicolorous or rarely bicolored, often pilose Antenna1 style with basal 2 annuli partly fused, appearing 3-annulate. Larger species, the wings with crossveins at least with weak clouds. The abdomen with three rows of white spots... a e... Stenotabanus (Phorcotabanus) Antenna1 style normal, clearly 4-annulate. Basal frontal callus usually with upper median prolongation; a median expanded callus and black hair patch often present. Frons generally narrower, seldom narrowed below, the basal callus frequently higher than wide, rareiy wider than high. Abdomen various, but not whitish and rarely with sublateral spots. Eyes rarely with more than 2 green bands Stenotabanus (Stenotabanus) Frons very broad, never over 1.5x as high as wide, with a transverse dark pollinose median band and callus wider than high. Wings with crossveins spotted and pattern of greyish nebulosity similar to species of Haematopota... Dasvbasis (Haematopotina) Frons variable, without median band Wings hyaline or spotted, but without grey Haemotopota pattern Frons relatively narrow; callus narrower than frons and prolonged above in a ridge or spur; tubercle at vertex generally with vestiges of 3 ocelli. If callus is subquadrate with short spur, the abdomen has prominent pale median triangles (See also couplet 39) Dasybasis (Agelanius) Frons variable; basal callus usually broad and nearly always as wide as frons, without median upper spur, and without ocellibearing tubercle at vertex. If callus is narrower than frons, the abdomen is without prominent pale triangles i... Dasy basis (Dasy basis) Basicosta with few to numerous setae, but seldom with setae as dense as on adjoining Costa. Generaliy with one or more of the following characters: tubercle at vertex; bare areas on face; partly sclerotized labella of proboscis; long dorsal spine on plate of third antenna1 segment; first antenna1 segment subcylindrical; wings with extensive dark pattern; swollen tibiae..... I Basicosta with setae as dense as on adjoining Costa. Rarely with any of the above characters Basal plate with acute dorsal angle or long spine which may reach beyond first annulus of style. Frons seldom over 4x as high as wide. Frontal callus clavate. occasionally as wide as frons at base, usually narrower. Palpi rather long and stout, never very slender nor markedly inflated basally. Eyes often pilose. \Vings very rarely entirely hyaline, generally with a dark discal patch below stigma and usually with discal cell fenestrate. Basicosta rarely bare Dicladocera Basal plate with dorsal angle obtuse, or frons much narrower _ Wings with extensive dark pattern not consisting of spots on crossveins. If wings apparently unmarked, then thorax prominently striped or frons exceedingly narrow and callus thread-like Wings hyaline, +e?, or.rr;tl-+ w l&l. Jfi*L- UUIIX t%n++pv-m FULL&--III primarily of dark spots around crossveins Wings with an irregular dark pattern of variable extent which always leaves clear areas surrounding all crossveins and fork of third vein. Frons narrow, 7 or more times as high as wide. Palpi very slender. Eyes unicolorous, bronzy in life... e...,.. Philipotabanus (Pbilipotabanus) Wings with small dark area below stigma, but crossveins not surrounded by clear spots when within dark areas. Palpi in-

8 44 REVISTA PERUANA DE ENTOMOLOGIA Vol Aated, antennae broad; proboscis hardi> longer than palpi, the labella large. Eyes green or brick-red in life _... Pbilipotahams (hh7oltrhanus) All crossveins and ends of all longitudinal veins close to wing margin with large dark spots, sometimes confluent. Frons broad, not over 3x as high as wide, the callus rounded, pointed above, as wide as frons. Vertex with distinct tubercle. Palpi slender, flattened, shorter than long proboscis. Thorax prominently striped Spilotabanus Wings never as extensively spotted, never with distinct spots on ends of longitudinal veins. Palpi not flattened, proboscis short Frons less than 4x as high as wide, generally wider. Eyes usually densely pilose. (See also couplet 33)... Dasybasis (Agelanius) Frons more than 4x as high as wide, generally narrower. Eyes very rarely pilose. 40 Wings hyaline or evenly tinted, the costal cell often darker, but never with spots on crossveins or apical clouds. Frontal callus clavate or ridge-like. Abdomen black or brown, nearly always with transverse bands at least on fourth tergite, rarely otherwise. At least scutellum and often mesonotum pale pollinose and pale haired, generally contrasting with abdomen. Appendix on fork of third vein absent. Eyes unbanded, dark... Leucotabanus Wing with clouds on at least discal crossveins, often with apical infuscations. If entirely hyaline or tinted, then abdomen and thorax not as above. Often with appendix on fork of third vein. Eyes unbanded, green or bronze, or light with single dark stripe. Frontal callus variable. Rarely with basicosta bare... Stypommisa Eyes generally pilose in female, always in male. First antenna1 segment with dorsal anterior margin much produced, cap-like. Eyes of female with 2 green bands, often bare or very sparsely pilose. Tubercle at vertex usually low and rounded, sometimes h \YJ;llOC TTA h r-tacc,,plnc Attpn 9 _- cent... a.. I.....A. -AA. ALCAl clouded, the first posterior cell generally at least coarctate, often closed and petiolate. Mainly south temperate species Poeciloderas Eyes very rarely sparsely pilose in female, generally bare in male Vertex with a strong tubercle, often bearing vestiges of ocelli. Eyes unicolorous, greenish black. Frontal callus clavate or ridge-like. Abdomen plain or banded, never striped. At least scutellum and often mesonotum pale pollinose and pale-haired, generally contrasting with abdomen. (See also couplet 40)...*... Leucotabanus Vertex with at most a small denuded spot. Eyes variously patterned or unicolorous, bare Tabanus SPECIES LIST Subfamily PANGONIINAE Tribe PANGONIINI Genus Esenbeckia Rondani Subgenus Esenbeckia Rondani * Esenbeckia bassleri Wilkerson and Fairchild San Martin, Moyobamba region (CAS); Huanuco, Upper Rio Huallaga (AMNH). - - Esenbeckia bitriangulata Lutz and Castro Lima, Verrugas Canyon (Lutz and Castro 1935); Lima, Surco (GBF!. Esenbeckia cisandeana Wilkerson and Fairchild Arnazonas, Rio Marafion (CAS); Amazonas, Rio Santiago (FSCA); Huanuco, Upper Rio Huallaga (CAS, AMNH); Huinuco, Tingo Maria, Monson Valley (CAS). -I; Esenbeckia gracihs Kroeber Loreto, Pebas (Kroeber 1931b); Amazonas, Rio Maraiion; Loreto, Iquitos; Loreto, Iquitos. Rio Itaya (All Berlin). * Esenbeckia rnelanogaster Lutz and Castro Lima, Verrugas Canyon (Lutz and Castro 1935); La Libertad, Cartavio (AMNH): Lima, Matucana (Cornell); Arequipa, Aplao (Carrasco); Arequipa, Aplao (GBF); La Libertad, Cartavio (GBF, San Diego). * Esenbeckia ni riventris Kroeber (Lima (Kroeber 193 lb); Peru (holotype in BMNH seen by GBF); Lima, Matucana (USNM). - - Esenbeckia osornoi quianense Fairchild Madre de Dios, Avispas (GBF). * Esenbeckia osornoi osomoi Fairchild Pasco, Pichis (Wilkerson and Fairchild XK_J J_ n: A_.:_. ^ If-hlf-\ 17031; lvlaulc UC UIUS, Avlsp s \LlYLI. - - Esenbeckia pecbumani Wilkerson and Fairchild Cuzco, Quincemil (CNC, GBF). - - Esenbeckia testaceiventris (Macquart ) Cuzco, Quincemil (LLP, GBF); Type labelled Quito, Perou in BMNH (GBF). - - Esenbeckia tigrina Wilkerson Huanuco, Cochicote (RCW, GBF). 1 Species of which we have actually seen specimens bear an asterisk ( 1. Those occurring in the Tambopata Reserve bear two asterisks (**).

9 Diciembre 1984 WILKERSON & FAIRCHILD: TABANIDAE OF TAMBOPATA 45 Esenbeckia translucens (Macquart ) E. Peru (Soukup 1943). Esenbeckia vulpes (Wiedemann) Loreto or Ucayali; Ucayali (Soukup 1945 ). - - Esenbeckia wygodzimky W ilkerson and Fairchild Cuzco, Machu Picchu (AMNH). Esenbeckia xanthoskela Wilkerson and Fairchild Amazonas, Bagua, Montenegro (AMNH, GBF); Huanuco, Tingo Maria (AMNH. GBF). Subgenus Proboscoides Philip. Esenbeckia armata (Williston) Pasco, Pichis (Soukup 1945). Esenbeckia ecuadorensis Lutz and Castro Avispas, Cuzco (determined for LLP by GBF, are probable E. Cl.) suturalis Rondani). - Esenbeckia fairchldi Philip Huanuco, Ytirac (CAS); Huhnuco, 67 mi. E. Tingo Maria (GBF). - : Esenbeckia rostrum Philip Loreto, Middle Rio Ucayali (AMNH); Avispas, Cuzco (determined by GBF for CNC, possibly misidentification for E. Cl.) suturalis). - - Esenbeckia schliugeri Philip HuBinuco, Tingo Maria, Monson Valley (CAS, GBF); Ucayali, Huallaga, Agua_ytia (GBF). ; +E sen b cc k za suturalis (Rondani) Madre de Dios, Avispas (LLP, GBF); Loreto, Middle Rio Ucayali (CAS). Tribe SCIONINI Genus Scuptia Walker Subgenus Scaptia Walker Scaptia aureonigra (Kroeber) Junm, Chanchamayo (Kroeber 193 IdI. Pasco and Jut-&r, Pichis and Pereni (USNM ). Scaptia rubriventris (Kroeber I Puno, vie. Limbani (GBF); Puno, Agualani, Limbani (GBF); Huancabamba(sic); Agualani, Limbani, Puno (C arrasco 1972 ); Cuzco, Rio Urubamba (USNM 1...$<sptz6 I il/u ~Lvla~qii%?) :m. /IAli Lima (Macquart 1838, probably in error, all other records from Chile). Subgenus Pseudoscione Lutz Scaptia albfions (Macquart) Pet-6 (Kroeber 1951, probably in error, no members of this subgenus known from Peru). Scaptza htipalpis (Macquart) Peti (Soukup 1945, probably in error, no members of this subgenus known from Peru). Genus Fidena Subgenus Fidena Walker - Fidena atripes (Riider) Peru: Madre de Dios, Cuzco, Arequipa, Peru (Carrasco 1972). Lima, Matucana; Lima, Surco; Arequipa, Lomas de CGpac nr. Chala; La Libertad, Cerro Campana, Trujillo (all GBF); Amazonas, Cerro Calla-Calla, 45 km E Balsas (CAS); Lima, Tambo de Viso; Cajamarca, Cascas; Cuzco, Sacsayhuamhn (MUSM). - - Fidena eriomeroides (Lutz) HuBnuco, Tingo Maria (Philip 1958, as F. nitida Philip); Pasco, Pozuzo (GBF). ; ;= - Fidena freemani Barrett Fidena howardi Fairchild Peru (GBF in CNC, locality doubtful as it is known elsewhere only from Panamh and Costa Rica). Fidena nigripes (Roder) Limatambo, Cuzco, (Carrasco 1972, probably determined by GBF). - : Fidena ochrapogon Wilkerson 1979 La Libertad, Cerro Campana, vie. Trujillo (GBF). + Fidena rhinophora (Bellardi) cuzco, Quincemil (GBF, CNC); Junin, Chanchamayo (GBF); Pasco and/or Junin, Pichis and Perene Valleys (Columbus). Fidena silvatica Brethes) Junin, Chanchamayo, vie. Rio Perene (Brethes 1920). * Fidena zonafis Kroeber Cuzco, Rio Urubamba, Umuhankiali (Kroeber 193 1~); Urubamba,... Satipo, Prov. Jauja, (Carrasco 1972); Huanuco, Cochicote; Madre de Dios, Avispas (both GBF); J unit-r, Chanchamavo (San Diego, CNC); Madre de Dios, Avispas; Cuzco, Quincemil (both CNC). Subgenus Laphriomyia Lutz Fidena kroeberi Fairchild nom. nov. for bicolor Kroeber Fidena mi ra b I I IS rufopilosus (Ricardo) Vergne toll., (Paris Museum). This identification not confirmed. Genus Scione Walker * Scione acer Philip Pasco, Osapampa (Philip 19% and in MUSM); Huinuco, Huanuco (Philip 1958); Peru ; Junin, Chanchamayo; Junin, Satipo (GBF, CNC, LLP). Scione albohirta Kroeber Cuzco, Urubamba (Kroeber 193Ob). Scione brevistriga Enderlein Madre de Dios, Pinipini (Soukup 19-15).

10 46 REVISTA PERUANA DE ENTOMOLOGIA Vol. 27 Scione cingulata (Enderlein) Cuzco, Callanga, (Enderlein 1925 ). - : Scione claripennis Ricardo Cuzco, Callanga (Ricardo 1900); Pasco, Pichis (Soukup 1945); Junin, Satipo (Cornell). Snbne crassa Szilady Cuzco, Hda. Cajon (Szilady 1926). * Scione equivexans Wilkerson Amazonas, 5 km N Pomacochas on rd. to Rioja (CAS). Scione fumzjpennis Kroeber Peru (Kroeber 193Ob). ; Scione fulva Ricardo Peru (Ricardo 1902); Cuzco, Laristal (Lares Valley) (Kroeber 1951); Cuzco, Laristal (BMNH). Scione fusca Ricardo Pasco, Huancabamba (Fairchild 1967); Huancabamba... Peru; Cuzco, Callanga, Pichis,... Cordillera, Peru. (Carrasco 1972 ). Scione grandis Philip Pasco, Rio Pionia (probably Rio Pichis, fide G. Lamas, pers. comm.), Puerto Bermtidez (Philip 1943). Scione huancabambae Kroeber Pasco, Huancabamba (Kroeber 1930b). Scime incompleta (Macquart) Peru: Rioja, Laristal,... Madre de Dies. (Carrasco 1972). This species probably = S. incompleta Kroeber, not Macquart, which = S. acer Philip. Scione Zimbativena Enderlein Norte de1 Peru. Huancabamba, (Carrasco 1972). Scione aurulans lurida Enderlein Peru: Laristal, (Carrasco 1972). Scione Zongirostris Brethes Junin, Chanchamayo (Brethes 1920). Scione obscurefemorata Kroeber Peru (Kroeber 193Ob). Scibne mfescens (Ricardo) Cuzco, CaIlanga (Ricardo 1900); Cuzco, Marcapata (Carrasco 1972, determined by GBF). Scione strigata (Enderlein) Genus Pityocera Giglio-Tos u ge?us CL Y*uyIJL Yu Ff hajl, YLLYI IL-A Pityocera cervus (Wiedemann) Peru: Pichis, Puerto Yessup. (Carrasco 1972). Cuzco, Quincemil (GBF); Madre de Di- OS, Avispas; Cuzco, Quincemil (CNC). Subfamily CHRYSOPSINAE Tribe CHRYSOPSINI Genus Chysops Cbysops boliviensis Kroeber Pasco, Oxapampa; Loreto, Iquitos (both MCZ). Chysops bulbicornis Lutz Loreto, Iquitos (Carrasco 1972). - Cbrssops calogaster Schiner Huanuco, Tingo Maria (Philip 1960). s:i: Chrysops emadorensis Lutz Loreto, Iquitos (Carrasco 1972). : Ch~sops incisus Macquart Loreto, Huallaga, Yurimaguas (Kroeber 1929b, as terzuifasciatus); Loreto, Iquitos and Amazonas (Carrasco 1972) Pichis, Cuzco. Ca- Uanga, Cajon (Carrasco 1972); Loreto, Iquitos (USNM). ;,- * Chrysops leucospihs Wiedemann Peru: Lares,... Madre de Dios, Pichis, Pto. Yessup, Vilcanota,... Tingo Maria. (Carrasco 1972); Huanuco, Tingo Maria, Monson Valley (GBF). Cb~sops hzi Kroeber Peru is given in original description but we doubt that it occurs in Peru since it is otherwise restricted to SE Brazil (Fairchild 19666). Ch~sops molestus Wiedemann Cuzco, Rosalina (Kroeber 1951); Madre de Dios, Avispas (RCW in CNC). Cb?~~sops peruvianus Kroeber Pachitea-Miindung; Pasco, Puerto Yessup; Pasco, Pichis; Cuzco, Callanga in Piiiipifiital (Pifiipini Valley) (all Munich) (Fairchild 1966d). ; : : Chysops varians var tardtls Wiedemann (C. Zaeta sublaeta Philip). Pilcopata, Cuzco (Carrasco 1972); Putumayo, El Encanto (error, Colombia) (Carrasco 1972); Ucayali, Unini (Carrasco 1972); Huanuco, Tingo Maria (Philip 1960); Cuzco, Quincemil (GBF, CNC); Huanuco, Tingo Maria (GBF, CAS, USNM, BMNH; Cuzco, PiIcopata (GBF); Madre de Dios, Avispas (CNC). + Chysops variegata (De Geer) Puerto Bermtidez, Pichis (Carrasco 1972); Coat-y am Amazonas Sivia (error, Coary, Amazonas is in Brazil and Sivia in Dept. Ayacucho, fide G. Lamas pers. comm.) (Carrasco 1972): Cuzco. Laristal (Carrasco 1972): Madre de Dies, Avispas (GBF); Loreto., Iquitos (US*N*M). 5:-z Cbrysops webevi Bequaert 1946 and species aff. weberi. Subfamily TABANINAE Tribe DIACHLORINI Genus Dasybasis Macquart Subgenus Dasybasis Macquart Dasybasis adornata (Kroeber) Cuzco, Paso de Lares (Coscaron and Philip, 1967). Qasvbasis albotibialis (Kroeber) 1931.

11 Diciembre _ 1984 WILKERSON & FAIRCHILD: TABANIDAE OF TAMBOPATA 47 Puno, Lake Titicaca (Coscaron and Philip, 1967). : ~a~~~basir barbata Coscaron and Philip, Puno, Limbani (GBF). Dasybasis bulbiscapens Coscaron and Philip Arequipa (Coscaron and Philip 1967, USNM). Dasybasis chilensis (Macquart) Lima, Miraflores (Kroeber 1951). Dasybasis cohmbiana (Enderlein) Huancabamba, Huancana and Cuzco, Marcapata; Apurimac, M6mara (both Carrasco 1972). ; ; L)asy;ybasis fairchildi Coscaron and Philip Puno, Tincopalca, 0. de Puno (Coscaron and Philip 1967); Ayacucho, Ayacucho (GBF). Dasybasis frequens (Kroeber) Cuzco (Coscaron and Philip 196-i); Puno, Caccackara, SE de Puno (Coscaron and Philip 1967); Puno, vie Limbani (Coscaron and Philip 1967, GBF); Peru, Cuzco, 4200 m. (BMNH). i: Dasybasis geminata Coscaron and Philip Puno, Desaguadero Lago Titicaca (Coscaron and Philip 1967, GBF, MUSM; Puno, Caccackara, SE de Puno (Coscaron and Philip 1967). Dasybasis inata Coscaran and Philip Puno, Imata; Puno, Caccackara SE de Puno; Lima (Coscaron and Philip 1967, GBF). Dasybasis kroeberi Coscaron and Philip Yura, Arequipa (both Coscaron and Philip 1967 ). Da$ash Zimbativena (Kroeber) uzco ; Madre de Dios ; Puno; Apurimac, Mhmara; Cuzco, Marcapata (all Coscaron and Philip 1967 ). Dasybasis neogrisescens (Kroeber) Mamara y Cuzco ( Coscaron and Philip 1967); Yacuacocha nr. Chiquisa; Junin, Tarma tboth GBF) ; J unm, Hda. Yanamarca near Jauja (MUSM); Cuzco, Sacsyahuaman (MUSM). Dusybasis penai Coscaron and Philip Puno, Limbani (GBF). - Dasybusis punensis (Hine) 1920.?Carnacani (Coscaron and Philip 1967); Arequipa?, Saracocha, bet\\wn Arequipa and Juliaca (GBF and Coscaron and Philip 1967); Puno, Lago Sarococha (GBF, MUSM); Lima, Ricripampa, Matucana (GBF); Puno, Lagunillas, near Puno (MUSM). Dasybasis shineri (Kroeber) Pachitea, Urubamba (Carrasco 1972). Dasybasis scbnusei (Kroeber) Cuzco and Sicuani (Coscaron and Philip 1967). Subgenus Agelanim Rondani Huaraz (GBF); Lima, Tambo de Visa (GBF, MUSM); Lima. Santa Eulalia \ alley (MUSAl); Lima, Rio Blanc0 (GBF, hlusm). : Daqbasis lauta var. sublauta Coscaron and Philip Lima, Surco; Lima, Verrugas Canyon (both Coscaron and Philip 1967). Subgenus Haematopotilza Coscaron and Philip Dasybasis pechmani Coscaron and Philip Lima (Coscaron and Philip 1967). Genus Styphocera Philip 5: Styphocera antelznina Philip ( 1969). Huanuco (Philip 1969). Genus Stenotabanus Lutz Subgenus Stenotabanus Lutz :+ Stenotabanus albilineavis Philip Huanuco, Tingo Maria, Monson Valley (Philip 1?60); Huinuco, Tingo Maria (GBF, AMNH); Madre de Dios, Avispas (GBF). : Stenotabunus bruesi (Hine) Lima, San Bartolome (Hine 1920, GBF); Cajamarca, Rio Namballe (GBF) Stenotabanus incipiens (Walker) Junin, Chanchamayo (Carrasco 1972 ); Huanuco, Yurac nr. Tingo Maria and Tingo Maria (Philip 1960); Madre de Dios, Avispas (GBF). - Stenotabanus obscures Kroeber Huinuco, Tingo Maria (Philip 1960, GBF, CAS, AMNH); Kosiiipata, Cuzco Carrasco 1972); Cuzco, QuincemiJ fgbf, CNC). - - Stenotabanus obscurm.flavofemoratus Kroeber Cuzco, Rio Urubamba (Kroeber 1929a); Hacienda Huayuri (Carrasco 1972); Cajamarca, Rio Charape (GBF): Huanuco, Tingo Maria (GBF, CliS); 1 Iuhnuco, Las Pal%as; _?Lima or Loreto, Cerro Azul (CAS). i: Stenotubunus pullidicornis Kroeber Huinuco, Tingo Maria; Junin, Valle Chanchamayo (GBF). * Stt?~iO&itii?dS pmiviensis Kroeber i929. Rio Pachitea (Kroeber 1929a); Urubamba and Umankilia (Umahuankiali, G. Lamas, pers. comm.) (Carrasco 1972); Cuzco, Quince- mil (GBF, Cornell, CNC); Madre de Dios, Loramayo (GBF); Loreto, Rio Ucayali (AMNH). Stenotabanus taeniotes (Wiedemann) Huinuco, Tingo Maria (Philip 1960). * Dasybasis lauta (Hine) Subgenus Phorcotabanus Fairchild Lima, Matucana; Lima, Surco (both Coscaran and Philip 1967, GBF, MUSM); Ancash, Stenotubanus cinereus (Wiedemann) 1821.

12 48 REVtSTA PERUANA DE ENTOMOLOGtA Vol. 27 Genus Himantostshs - Hilz7antost~lzrs intermediur Lutz Puno, Yahuarmayo (Lutz 1913 ): Cuzco, Quincemil (Carrasco 1972); Cuzco, Quincemil; Madre de Dios (both CNC). Genus Diacblorur Lutz 5: Diacblorus bicinctus (Fabricius) ; Dracblorus bzmaculatus ( Wiedcmann) San Martin, hfoyobamba (CAS). : : Diacbiorus curvipes (Fabricius) Pucallpa; Pachtea (Krocber 195 I ); Cuzco, Quincemil; (GBF, Cornell). : Diacblorus fusczstigma Lutz Loreto, Rio Nanay (GBF). i: Diacblorus heppneri VCrilkerson and Fairchild : Diacblorus nuneztovari Fairchild and Ortiz Diacblorus pecbumarzi Fairchild cuzco, Quincemil; Loreto, Iquitos t both Fairchild 1972 ). Cc Diacblorus scutellatus (Macquart) Diacblorus xynus Fairchild Cuzco, Quincemil (Fairchild Cornell). Genus Bolbodimyia Bigot Bolbodiwyia bicolor Bigot Junin, vie. la Merced (CAS). 5: Bolbodiwyia celeroides Stone Cuzco, Quincemil (GBF, CNC). f: Bolbodimyia desecta Enderlein Junin, Chanchamayo Valley (Stone 1954); Cuzco, Quincemil (CNC). : Bolbodimyia nigra Stone Ucayali, Cinchona (Stone 1954); CUZCO, Quincemil (CNC 1. Genus Cblorotabanus Lutz i i : Cblorotabanus inanis (Fabricius) Ucayali, Santa Sofia, Rio Utoquinia (GBF, MUSM). ; : Cblorotabanus parviceps (Kroeber) Huanuco, Tingo Maria (Philip and Fairchild 1956); Kosiiipata, Cuzco (Carrasco 1972, determined by GBF) Cblorotabanus?leucocblorus Fairchild. Genus Pbaeotabanus Lutz Pbaeotabanus apbanopterus (Wiedemann) Ucayali, Rio Ucayali (Philip 1960). : Pbaeotabanus cajennensis (Fabricius) Peru (Kroeber 1951). ; - Pbaeotabanus fervens (Linnaeus San Martin, Navarro (MUSM). Pbaeotabanus in0 tescens (Walker) 1854 (CAS). Pbaeotabams limpidapex (Wiedemann J lunin, Chanchamayo (Ktoeber 1930~ probably error). Pbaeotabanus nigriffavur Kroeber Ucayali, Pucallpa; Ucayali, Rio Ucayali; Huanuco, Tingo Maria (all Philip 1960); Ucayali, 20 mi. W Pucallpa (CAS). Pbaeotaballus pbaropterus Fairchild Madre de Dios, Avispas (GBF, CNC). Genus Sp,;kttabams Fairchild Spilotabanus multiguttatus (Kroeber ) Machu Picchu, Cuzco (Carrasco 1972 ); Huinuco, Carpish (GBF). Genus Eutabanus Kroeber Eutabanus pictus Kroeber Ucayali, NO Ucayali (Kroeber 1930a, BMNH). Genus Acanthocera Macyuart. Acantbocera bequarti Fairchild and Aitken Acantbocera fornzosa Kroeber Loreto, Iquitos (Kroeber 193Od). Acantbocera marginah Walker No localitv given (Carrasco 1972); Huanuco, Yurac, 67 mi E Tingo Maria (CAS); Madre de Dios, Avispas; Cuzco, Quincemil (both CNC). Genus Querbetia Fairchild - - Querbetia bequaerti Fairchild Huanuco, Tingo Maria (Fairchild MCZ). : Querbetia inopinatus Fairchild Madre de Dios, Avispas (Fairchild CNC). Genus Dicbelacera Macquart Subgenus Dicbelacera Macquart 1963, 1972, Dicbelacera cervicornis (Fabricius) Loreto, Pebas; Loreto, Iquitos (both Kroeber 1951). ; : Dicbelacera fuscipes Lutz Peru (Munich, possibly mislabeliedi. Dicbelacera marginata Macquart Loreto, Iquitos (Fairchild and Philip 1960); Loreto, Pebas (Kroeber 1951). Dicbelacera villavoensis Fairchild and Philip Huanuco, Tingo Maria, Monson Valley (Fairchild and Philip 1960); Loreto, Iquitos (GBF); Huanuco, Tingo Maria; Amazonas, Olmos-Marafion Highway, km 292, E of Montenegro; Cuzco, Quincemil; Madre de Dios, Avispas (CNC). i: Dicbelacera submarginata Lutz 1915.

13 Dlciembre 1984 WILKERSON & FAIRCHILD: TABANIDAE OF TAMBOPATA 49 Ucavali, Middle Ucavali; Loreto, Rio Mara&; Huanuco, Tingo Maria, Rio Hurtilaga; Huanuco, Tingo Maria, Monson Valley; San Martin, Roque: Loreto, Iquitos, Rio Itaya; Cajamarca, Rio Charape (all Fairchild and Philip 1960): Huanuco. Cochicote (USNM); Amazonas, Huambo (Kroeber 1930a); Ucapaii or Loreto, Middle Ucayali; Cuzco, Quincemil; Loreto, Iquitos; Madre de Dios, Avispas; Huanuco, Cueva de las Pavas (all GBF): Hu6nuco, 5mi SW Las Palmas (CAS); Cuzco, Quincemil; Madre de Dios, Avispas (both CNC); Loreto, Iquitos, Rio Itaya (GBF, ret. to Lund); Cuzco, Buenos Aires, 53km W Pilcopata, 2280m (USNM). Subgenus Idiochelacera Fairchild Dichelacera subcallosa Fairchild and Philip Huinuco, Cochicote (GBF). Genus Catachlor-ops Lutz Subgenus Catacblorops Lutz Catachlorops balteratus Kroeber A specimen in Paris from Surinam was part of Macquart s type series of Zuctuosa. but a different species, and agreed with one female, Iquitos, Loreto, Peru, Shannon COIL, det. halteratus Kroeber by Bequaert. Halteratus described from Guianas. Kroeber s type is in Vienna. Subgenus Amphicblorops Lutz Catacblorops uespertinus (Bequaert and Renjifo) Huambo, Rosalina, Urubamba (Carrasco 1972); Peru (Paris); Cuzco, Quincemil (CNC). Subgenus Hadrocblorops Fairchild Catacblorops scutellatus (Macquart) Peru (Philip 1960). Subgenus Psalidia Enderlein he Catacblorops rubiginosus (Summers) Genus Dasycbela Enderlein Subgenus Dasycbela Enderlein Subgenus Triceratoqia Bequaert Daswbela birarnula Fairchild Tunin, La Merced, Hacienda La Salud (Leiden Museum); Junin, Chanchamapo tgbf ret. to Leclercq I. Daqchela macilrtyrei Bequaert Cuzco. Quincemil (GBF, CNC). Genus Diclndocera Lutz Dicladocera bellicosa (Brethes) Junin, Chanchamayo and Chanchamayo, La Merced (GBF). Dicladocera castanea (Barreto) Hutinuco (Barreto 1949). Junin, Valle Chanchamayo; Cuzco, Quincemil (GBF, Cornell); Huanuco, Cochicote ( USNrM). ; Dicladocera clams!schiner) Junm, Huacapistana, Rio Tarma; Lima, Verrugas (GBF). Dicladocera fulvicornis Kroeber Peru, Pichis (Kroeber 1931, Fairchild 1975). 5: Dicladocera griseipennis Kroeber Madre de Dios (Fairchild 1966). * Dicladocera boppi Enderlein Arequipa, Carcani (Enderlein 1927); Arequipa ; Arequipa, nr. Arequipa (both GBF, Paris). Dicladocera macula (Macquart) Loreto, Rio Huallaga; Lima, Matucana (both GBF). + DicZadocera maculistigma Enderlein Lima (Enderlein 1925); Cuzco, Limatambo (Carrasco 1972); Lima, Santa Eulalia (USNM). DtLfy mines (Schiner) 1868., Huancabamba (Kroeber 1951); Peru (Paris). Dicladocera nigrocoerulea (Rondani) Pasco, Pichis (Kroeber 1931). A doubtful record, as no Peruvian specimens determined by Kroeber seen. Paratypes are from Colombia, Venezuela, Brasil. Dicladocera neosubmacula Kroeber Huanuco. Tingo Maria (Philip 1960); Cuzco, Quince&l (CNC, Cornell). A - Dicladocera nova Kroeber Cuzco, Quincemil; Cuzco, Valle de1 Rio Lares (GBF, Cornell, Dresden Museum and CNC). ; Dasycbela amazonensis (Barreto) Genus Stibasoma Schiner _ * n.i n I. Loreto ( GBF). Subgenus :, tzbasonza Schmer Dasycbela inca (Philip) ir Stibasoma jlatliventre (Macquart) Huanuco, Tingo Maria (Philip 1960, GBF); Loreto, Iquitos, Rio Itaya (Berlin Museum). Junin, Satipo; Ucayali, Cinchona (both USNM); Junin, Region de Perene (Paris). Genus Clyptotylus Lutz - Dasycbeia peruviana (Bigot) : Cryptotylus unicolor (Wiedemann) Peru (Bigot 1892 );?lgnapo (Kroeber Peru (Kroeber 1951); Ucayali, Santa Sofia, 1951); Peru (BMNH). Rio Utoquinia (GBF, MUSM).