CONTRIBUTION TO THE BUTTERFLY FAUNA (HESPERIOIDEA & PAPILIONOIDEA) OF THE VELEBIT MOUNTAIN, CROATIA

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1 NAT. CROAT. VOL. 16 No ZAGREB March 31, 2007 original scientific paper/izvorni znanstveni rad CONTRIBUTION TO THE BUTTERFLY FAUNA (HESPERIOIDEA & PAPILIONOIDEA) OF THE VELEBIT MOUNTAIN, CROATIA IVA MIHOCI, MARTINA [A[I] &MARIJANA VUKOVI] Croatian Natural History Museum, Department of Zoology Demetrova 1, HR Zagreb, Croatia Mihoci, I., [a{i}, M. & Vukovi}, M.: Contribution to the butterfly fauna (Hesperioidea & Papilionoidea) of the Velebit Mountain, Croatia. Nat. Croat., Vol. 16, No 1., 29 62, 2007, Zagreb. During the years 2005 and 2006, 106 butterfly species were identified on Mt. Velebit. The estimated distribution, diversity and biogeographical characteristics of each species are presented in this paper as well as a comparison between recently estimated butterfly fauna and that identified in the past century. Current check-list of 137 butterflies from Mt. Velebit is contained in the Appendix. Key words: butterflies, fauna, distribution, diversity, protection, Mt. Velebit Mihoci, I., [a{i}, M. & Vukovi}, M.: Prilog poznavanju danjih leptira (Hesperioidea & Papilionoidea) Velebita, Hrvatska. Nat. Croat., Vol. 16, No 1., 29 62, 2007, Zagreb. U dvogodi{njem razdoblju (2005. i 2006.) na prostoru Velebita zabilje`eno je 106 vrsta danjih leptira. U radu se navodi njihovo rasprostranjenje, raznolikost kao i biogeografska karakteristika za svaku utvr enu vrstu. Isto tako uspore uje se fauna danjih leptira toga prostora utvr ena u pro{lom stolje}u s recentnim podacima, te se na temelju toga daje popis 137 vrsta danjih leptira Velebita. Klju~ne rije~i: leptiri, fauna, rasprostranjenost, raznolikost, za{tita, Velebit INTRODUCTION With a length of 145 km, Mt. Velebit is the longest mountain of the Dinaric Alps. It spreads from the Vratnik pass above Senj in the north-west to the Zrmanja River in the south-east. Average distance from the coastal area to the Lika continental bed is 14 kilometres. The coastal slope of the mountain is much steeper, distinctly craggy and bare with two distinct longitudinal terraces. The continental slope is much gentler but can be abrupt, mostly forested and overgrown in vegetation, mostly without terraces, but with deeply truncated valleys (FORENBACHER, 2001). Of an area of 2274 km 2 only 41 km 2 refers to the vertical zone above 1600 m a.s.l. (POLJAK, 1974). In all, the important characteristics of Mt. Velebit are its low indentedness, abrupt slopes, significant daily temperature variations, and mixed continental, Mediter- Croatian Natural History Museum, Demetrova 1, Zagreb, Croatia

2 30 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia ranean and mountainous climate regimes and a combination of central European and Mediterranean vegetation. On Mt. Velebit different ecological and climatic characteristics have conditioned high biotope diversity which has, respectively, ensured high faunistic, and hence butterfly diversity. According to KU^INI] et al. (1995) the butterfly fauna of the Mt. Velebit comprises 115 species. This number is based on the analysis of entomological collections from the Croatian National History Museum (CNHM) in Zagreb and all the existing literature referring to the butterfly fauna of Mt. Velebit. The goal of this paper is to contribute to the knowledge about the butterfly diversity of Mt. Velebit and to supply a new butterfly check-list for Mt. Velebit. MATERIAL AND METHODS Butterflies were collected with an entomological net, observed and photographed in the years 2005 and 2006 at 70 localities on Mt. Velebit (Fig. 1). All collected Fig. 1. Researched area Mt Velebit, Croatia

3 Nat. Croat. Vol. 16(1), Tab. 1. Alphabetical list of researched locations, belonging to the Sjeverni Velebit National Park (*), the Velebit Nature Park (**) and the Paklenica National Park (***), altitude of each locality and number of found species on each locality (coordinates are given for unprecisely defined localities). Ordinal number Location Number of found species Altitude (m a.s.l.) 1. above Borisov dom mt. hut *** above Dundovi} Pade`* ( / ) after Bru{ane (direction B. O{tarije)** Ani}a Luka*** Apati{an** Babi} Si~a* Babino jezero*** Babrova~a* Ba~i} Duliba** Bakovac** Ba{ke O{tarije** Borisov dom mt. hut*** Bru{ane** Borova~ki Pade`** Buljma (pass)*** Bunovac*** Bu`im** ^a~i} Draga** Cigani{te* Dabarska kosa** Duplje* forest hut Paklenica NP*** from Begova~a to Krasno** Glamo~eva Duliba* Gornja Klada** Jadovno** Jazmaku{a** Jezera** Konjsko** Kosinjski Bakovac** Krasno** Kriva~a* Kri`i}i*

4 32 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia Ordinal number Location Number of found species Altitude (m a.s.l.) 34. Kubus** Ljubica creek (near Ba{ke O{tarije)** Mala Mo~ila*** Malovan (M. stanovi)*** Malovansko jezero*** Modri} dolac* Oltari** Pari}i*** Petra{ica** Polo`ine** Radu~** Rami}i*** Struge*** [arinac* [prenga** [tirova~a* Sun erac** [vica Te`akova~ko vrelo** to Bakovac** ( / ) to Bu`im** ( / ) to Dabarska kosa** ( / ) to Klementa** ( / ) to Kugina ku}a mt. hut* ( / ) to Mala Mo~ila*** ( / ) to Malovan (M. stanovi) *** ( / ) to Oltari** ( / ) to Sijasetska draga ** Trnovac** Tudorevo* turn to Pandore** ( / ) turn to Zavi`an mt. hut* ( / ) Velika Mo~ila*** Veliki Javornik*** Veliki Lubenovac* Vu~jak* Vodice*

5 Nat. Croat. Vol. 16(1), specimens are deposited in the butterfly collection of the Department of Zoology, CNHM, in Zagreb. Systematics follows KARSHOLT & RAZOWSKI (1996) and taxonomic determination regarding wing morphology was done according to TOLMAN & LEWINGTON (1997). Biogeographical analysis follows HRUBY (1964) and for species that were not listed there, it follows SIJARI] et al. (1984) and HIGGINS & RILEY (1993). The analysis of the morphological characteristics of the genitals of specimens from the Leptidea genus was done according to LORKOVI] (1993), JAK[I] (1998), LELO (2002) and LELO (2003). Altitudes were approximately determined using the GPS device Garmin etrex Summit and geographical sheet maps of Mt. Velebit (1:25000). RESULTS In the two year period we counted 106 species belonging to five butterfly families at 70 localities (Tab. 1). Species are listed according to the systematics by KARSHOLT & RAZOWSKI (1996) and the list of localities for each species is given in chronological order of the finding species at a particular finding site. In parentheses beside the species name, the number of stars indicates the presence of the species in the: * Northern Velebit National Park, ** Velebit Nature Park and *** Paklenica National Park. We have recorded 69 species for the Northern Velebit National Park, 96 species for the Velebit Nature Park and 55 species for the Paklenica National Park. Hesperioidea Hesperiidae (6 species) Pyrginae Erynnis tages (Linnaeus, 1758) (**): to Oltari, June 17 th, Carcharodus floccifera (Zeller, 1847) (*): Tudorevo, July 19 th, Hesperiinae Thymelicus lineola (Ochsenheimer, 1808) (* / **): Bu`im, July 7 th, 2006; Kosinjski Bakovac, July 19 th, Vodice, July 20 th, 2006; Jadovno, July 21 st, Thymelicus sylvestris (Poda, 1761) (* / ** / ***): Babi} Si~a, July 25 th, 2005; Dabarska kosa, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Jadovno, July 6 th, 2006; Bu`im, July 6 th, 2006; Ljubica, July 6 th, 2006; Bakovac, July 7 th, Vodice, July 20 th, 2006; [arinac, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Bunovac, July 26 th, Hesperia comma (Linnaeus, 1758) (* / Vodice, July 20 th, 2006; Dabarska kosa July, 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006.

6 34 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia Ochlodes venata (Bremer & Grey, 1853) (* / ** / ***): Dabarska kosa, June 18 th, 2005; Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Oltari, July 26 th, 2005; turn to Pandore, July 26 th, 2005; Apati{an, July 26 th, 2005; [tirova~a, July 27 th, 2005; [prenga, July 27 th, 2005; Ljubica, July 27 th, 2005; Sun erac, July 27 th, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; above Borisov dom mt. hut, August 26 th, 2005; Trnovac, June 20 th, 2006; Jadovno, June 23 rd, 2006; Ljubica, July 6 th, 2006; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; from Begova~a to Krasno, July 7 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Kri`i}i, July 20 th, 2006; Cigani{te, July 20 th, 2006; Babrova~a, July 20 th, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006; Pari}i, July 27 th, Papilionoidea Papilionidae (4 species) Parnassiinae Parnassiini Parnassius mnemosyne (Linnaeus, 1758) (* / **): Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; [tirova~a, June 18 th, 2005; Jezera, July 26 th, 2005; Tudorevo, July 19 th, 2006; Vu~jak, July 20 th, Parnassius apollo (Linnaeus, 1758) (** / ***): Polo`ine, July 27 th, 2005; Dabarska kosa, July 27 th, 2005; from Begova~a to Krasno, July 7 th, 2006; Dabarska kosa, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Jadovno, July 21 st, 2006; Struge, July 27 th, Papilioninae Iphiclides podalirius (Linnaeus, 1758) (* / ** / ***): to Oltari, June 17 th, 2005; turn to Pandore, June 17 th, 2005; Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; Konjsko, June 18 th, 2005; [vica, July 25 th, 2005; Dabarska kosa, July 27 th, 2005; Kubus, July 28 th, 2005; Ani}a Luka, August 1 st, 2005; Babrova~a, July 20 th, 2006; forest hut Paklenica NP, July 27 th, Papilio machaon Linnaeus, 1758 (** / ***): [vica, July 25 th, 2005; Jezera, July 26 th, 2005; Ljubica, July 27 th, 2005; Mala Mo~ila, August 2 nd, 2005; to Bu`im, July 6 th, 2006; to Malovan (M. stanovi), July 26 th, 2006; Malovan (M. stanovi), July 26 th, Pieridae (11 species) Dismorphiinae Leptidea sinapis (Linnaeus, 1758) (* / ** / ***): Borova~ki Pade`, June 18 th, 2005; [tirova~a, June 19 th, 2005; [vica, July 25 th, 2005; Oltari, July 26 th, 2005; turn to Pandore, July 26 th, 2005; Apati{an, July 26 th, 2005; Ljubica, July 27 th, 2005; to Bu`im, July 6 th, 2006; Bru{ane, July 6 th, 2006; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Veliki Lubenovac, July 19 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Jadovno, July 21 st, 2006; Malovan (M. stanovi), July 26 th, 2006; Pari}i, July 27 th, 2006; Jadovno, July 28 th, 2006; Radu~, July 28 th, 2006; Bunovac, July 28 th, 2006.

7 Nat. Croat. Vol. 16(1), Pierinae Antocharini Antocharis cardamines (Linnaeus, 1758) (* / **): Borova~ki Pade`, June 18 th, 2005; Babi} Si~a, June 17 th, 2005; to Dabarska kosa, June 18 th, Pierini Aporia crataegi (Linnaeus, 1758) (* / **): [vica, June 16 th, 2005; turn to Pandore, June 17 th, 2005; to Oltari, June 17 th, 2005; Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; Apati{an, July 26 th, 2005; Sun erac, July 28 th, 2005; Trnovac, June 20 th, 2006; Trnovac, June 23 rd, 2006; Jadovno, June 23 rd, 2006; Jadovno, July 6 th, 2006; to Bu`im, July 6 th, 2006; Bu`im, July 6 th, 2006; Ljubica, July 6 th, 2006; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; to Klementa, July 7 th, 2006; Petra{ica, July 7 th, 2006; [tirova~a, July 7 th, 2006; Veliki Lubenovac, July 19 th, 2006; Glamo- ~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; Ba~i} Duliba, July 20 th, Pieris brassicae (Linnaeus, 1758) (* / ** / ***): to Oltari, June 17 th, 2005; Ljubica, June 18 th, 2005; Sun erac, July 28 th, 2005; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Veliki Lubenovac, July 19 th, 2006; Glamo~eva Duliba, July 20 th, 2006; to Sijasetska draga, July 26 th, 2006; Malovan (M. stanovi), July 26 th, Pieris mannii (Mayer, 1851) (* / ** / ***): Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Konjsko, June 18 th, 2005; [vica, July 25 th, 2005; Babi} Si~a, July 25 th, 2005; Oltari, July 26 th, 2005; Apati{an, July 26 th, 2005; [tirova~a, July 27 th, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; above Borisov dom mt. hut, August 26 th, 2005; Pari}i, August 26 th, 2005; Rami}i, August 26 th, 2005; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Petra{ica, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Veliki Lubenovac, July 19 th, 2006; Vu~jak, July 20 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Duplje, July 20 th, 2006; [arinac, July 20 th, 2006; Babrova~a, July 20 th, 2006; Ba~i} Duliba, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; to Malovan (M. stanovi), July 26 th, 2006; Malovan (M. stanovi), July 26 th, 2006; Ani}a Luka, July 27 th, 2006; forest hut Paklenica NP, July 27 th, 2006; Borisov dom mt. hut, July 27 th, 2006; Jadovno, July 28 th, 2006; Bunovac, July 28 th, Pieris rapae (Linnaeus, 1758) (* / ** / ***): Babi} Si~a, July 25 th, 2005; Oltari, July 26 th, 2005; Apati{an, July 26 th, 2005; Polo`ine, July 27 th, 2005; Dabarska kosa, July 27 th, 2005; Ani}a Luka, August 1 st, 2005; Mala Mo~ila, August 2 nd, 2005; Dabarska kosa, July 21 st, 2006; Borisov dom mt. hut, July 27 th, Pieris ergane (Geyer, 1828) (* / ** / ***): Babi} Si~a, July 17 th, 2005; Dabarska kosa, June 18 th, 2005; Ani}a Luka, August 1 st,

8 36 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia 2005; Mala Mo~ila, August 2 nd, 2005; Rami}i, August 26 th, 2005, Trnovac, June 20 th, 2006; above Dundovi} Pade`, July 19 th, Vodice, July 20 th, Pieris napi (Linnaeus, 1758) (* / ** / ***): Babi} Si~a, June 17 th, 2005; to Dabarska kosa, June 18 th, 2005; [tirova~a, July 27 th, 2005; Ljubica, July 27 th, 2005; Velika Mo~ila, August 2 nd, 2005; Pari}i, August 26 th, 2005; Bu`im, July 7 th, 2006; [tirova~a, July 7 th, 2006; from Begova~a to Krasno, July 7 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, Coliadinae Colias croceus (Fourcroy 1785) (* / ** / ***): Dabarska kosa, June 18 th, 2005; [vica, July 25 th, 2005; turn to Pandore, July 26 th, 2005; Sun erac, July 28 th, 2005; Pari}i, August 26 th, 2005; Petra{ica, July 7 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Tudorevo, July 19 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; [arinac, July 20 th, 2006; to Sijasetska draga, July 26 th, 2006; Bunovac, July 26 th, 2006; Malovan (M. stanovi), July 26 th, 2006; Bunovac, July 28 th, Gonepteryx rhamni (Linnaeus, 1758) (* / ** / ***): Babi} Si~a, June 17 th, 2005; Gornja Klada, June 17 th, 2005; Dabarska kosa, June 18 th, 2005; [vica, July 25 th, 2005; Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Apati{an, July 26 th, 2005; [tirova~a, July 27 th, 2005; Ljubica, July 27 th, 2005; Sun erac, July 28 th, 2005; Mala Mo~ila, August 2 nd, 2005; Pari}i, August 26 th, 2005; Rami}i, August 26 th, 2005; Jadovno, July 6 th, 2006; Bru{ane, July 6 th, 2006; Kosinjski Bakovac, July 19 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006: to Malovan (M. stanovi), July 26 th, 2006; Malovan (M. stanovi), July 26 th, 2006; Radu~, July 28 th, Gonepteryx cleopatra (Linnaeus, 1767) (**): Dabarska kosa, June 18 th, 2005; Jezera, July 26 th, Lycaenidae (25 species) Riodininae Hamearis lucina (Linnaeus, 1758) (* / **): Babi} Si~a, June 17 th, 2005; to Oltari, June 17 th, 2005; Dabarska kosa, June 18 th, Lycaeninae Lycaenini Lycaena virgaureae (Linnaeus, 1758) (* / **): Borova~ki Pade`, June 18 th, 2005; Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; turn to Pandore, July 26 th, 2005; Apati{an, July 26 th, 2005; [tirova~a, July 27 th, 2005; Dabarska kosa, July 27 th, 2005; Ljubica, July 27 th, 2005; Sun erac, July 28 th, 2005; Jadovno, July 6 th, 2006; ^a~i} Draga, July 6 th, 2006; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Petra{ica, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; Jadovno, July 21 st, 2006; Jadovno, July 28 th, 2006.

9 Nat. Croat. Vol. 16(1), Lycaena alciphron (Rottemburg, 1775) (**): Ljubica, July 6 th, Lycaena candens (Herrich-Schäffer, 1844) (* / **): Jadovno, June 23 rd, 2006; Veliki Lubenovac, July 19 th, Eumaeini Satyrium w-album (Knoch, 1782) (*): Glamo~eva Duliba, July 20 th, Satyrium pruni (Linnaeus, 1758) (**): Trnovac, June 20 th, Satyrium spini (Denis & Schiffermüller, 1775) (* / **): to Dabarska kosa, June 18 th, 2005; Babi} Si~a, July 25 th, 2005; Kosinjski Bakovac, July 19 th, 2006; Tudorevo, July 19 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; [arinac, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, Satyrium ilicis (Esper, 1779) (**): Ba{ke O{tarije, July 7 th, Polyommatini Cupido minimus (Fuessly 1775) (* / **): Kri`i}i, July 20 th, 2006; Glamo~eva Duliba, July 20 th, 2006; [arinac, July 20 th, 2006; Ba~i} Duliba, July 21 st, 2006; Jadovno, July 21 st, Cupido alcetas (Hoffmannsegg, 1804) (**): Bu`im, July 7 th, Celastrina argiolus (Linnaeus, 1758) (* / ** / ***): Vu~jak, July 25 th, 2005; Dabarska kosa, July 27 th, 2005; Ljubica, July 27 th, 2005; Sun erac, July 28 th, 2005; Mala Mo~ila, August 2 nd, 2005; above Borisov dom mt. hut, August 26 th, 2005; Pari}i, August 26 th, 2005; Dabarska kosa, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Pari}i, July 27 th, Scolitantides orion (Pallas, 1771) (** / ***): to Dabarska kosa, June 18 th, 2005; Pari}i, August 2 nd, Glaucopsyche alexis (Poda, 1761) (**): Ljubica, June 18 th, Maculinea arion (Linnaeus, 1758) (* / **): [vica, July 25 th, 2005; [tirova~a, July 27 th, 2005; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Jadovno, July 21 st, Maculinea rebeli (Hirschke, 1904) (**): Jadovno, July 6 th, 2006; Bakovac, July 7 th, 2006.

10 38 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia Plebeius argus (Linnaeus, 1758) (** / ***): [vica, June 16 th, 2005; Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; [vica, July 25 th, 2005; Velika Mo~ila, August 2 nd, 2005; Ljubica, July 6 th, 2006; Bu`im, July 7 th, 2006; Ba~i} Duliba, July 21 st, 2006; Jadovno, July 21 st, 2006; Bunovac, July 28 th, Plebeius idas (Linnaeus, 1761) (***): [vica, June 16 th, 2005; [vica, July 25 th, 2006; Velika Mo~ila, August 2 nd, Aricia agestis (Denis & Schiffermüller, 1775) (* / ** / ***): Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Oltari, July 26 th, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; Trnovac, June 23 rd, 2006; Jadovno, June 23 rd, 2006; to Bakovac, July 7 th, 2006; Petra{ica, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Kri`i}i, July 20 th, Vodice, July 20 th, 2006; [arinac, July 20 th, 2006; Babrova~a, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; Bunovac, July 26 th, 2006; Pari}i, July 26 th, 2006; Bunovac, July 28 th, Polyommatus semiargus (Rottemburg, 1775) (* / ** / ***): Apati{an, July 27 th, 2005; Trnovac, June 23 rd, 2006; Jadovno, June 23 rd, 2006; Jadovno, July 6 th, 2006; Ljubica, July 6 th, 2006; Bakovac, July 7 th, 2006; Petra{ica, July 7 th, 2006; Tudorevo, July 19 th, Vodice, July 20 th, 2006; Te`akova~ko vrelo, July 21 st, 2006; Malovan (M. stanovi), July 26 th, Polyommatus dorylas (Denis & Schiffermüller, 1775) (** / ***): Jezera, July 26 th, 2005; Ba~i} Duliba, July 21 st, 2006; to Sijasetska draga, July 26 th, 2006; Malovan (M. stanovi), July 26 th, 2006; Veliki Javornik, July 27 th, 2006; Struge, July 27 th, 2006; Ani}a Luka, July 27 th, Polyommatus amandus (Schneider, 1792) (* / ** / ***): Dabarska kosa, June 18 th, 2005; Jezera, July 26 th, 2005; Jadovno, July 6 th, 2006; Ljubica, July 6 th, 2006; Petra{ica, July 7 th, 2006; [tirova~a, July 7 th, 2006; Tudorevo, July 19 th, 2006; Veliki Lubenovac, July 19 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; [arinac, July 20 th, 2006; Ba~i} Duliba, July 21 st, 2006; Malovan (M. stanovi), July 26 th, 2006; Malovansko jezero, July 26 th, 2006; Bunovac, July 28 th, Polyommatus icarus (Rottemburg, 1775) (* / ** / ***): [vica, June 16 th, 2005; to Oltari, June 17 th, 2005; turn to Pandore, June 17 th, 2005; Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Borova~ki Pade`, June 18 th, 2005; Babi} Si~a, July 25 th, 2005; [tirova~a, July 27 th, 2005: Ljubica, July 27 th, 2005; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Petra{ica, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Tudorevo, July 19 th, Vodice, July 20 th, 2006; Babrova~a, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Jadovno, July 21 st, 2006; Malovan (M. stanovi), July 26 th, 2006; Borisov dom mt. hut, July 27 th, 2006; Pari}i, July 27 th, 2006; Radu~, July 28 th, 2006; Bunovac, July 28 th, 2006.

11 Nat. Croat. Vol. 16(1), Polyommatus daphnis (Denis & Schiffermüller, 1775) (* / ** / ***): Mala Mo~ila, August 2 nd, 2005; above Dundovi} Pade`, July 19 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; [arinac, July 20 th, 2006; Babrova~a, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Ani}a Luka, July 27 th, 2006; Pari}i, July 27 th, Polyommatus bellargus (Rottemburg, 1775) (* / ** / ***): to Oltari, June 17 th, 2005; turn to Pandore, June 17 th, 2005; Jezera, July 26 th, 2005; Apati{an, July 26 th, 2005; Pari}i, August 26 th, Vodice, July 20 th, 2006; Pari}i, July 27 th, Polyommatus coridon (Poda, 1761) (* / ** / ***): Dabarska kosa, July 27 th, 2005; Ljubica, July 27 th, 2005; Ani}a Luka, August 1 st, 2005; Mala Mo~ila, August 26 th, 2005; Pari}i, August 26 th, 2005; Rami}i, August 26 th, 2005; Tudorevo, July 19 th, Vodice, July 20 th, 2006; [arinac, July 20 th, 2006; Babrova~a, July 20 th, 2006; Duplje, July 20 th, 2006; Jadovno, July 21 st, 2006; Malovan (M. stanovi), July 26 th, 2006; Veliki Javornik, July 27 th, 2006; Buljma (pass), July 27 th, 2006; Ani}a Luka, July 27 th, 2006; Jadovno, July 28 th, 2006; Radu~, July 28 th, 2006; Bunovac, July 28 th, Nymphalidae (60 species) Libytheinae Libythea celtis (Laicharting, 1782) (** / ***): Konjsko, June 18 th, 2005; Rami}i, August 26 th, 2005; Te`akova~ko vrelo, July 21 st, Heliconiinae Argynnis paphia (Linnaeus, 1758) (* / ** / ***): Gornja Klada, June 17 th, 2005; [vica, July 25 th, 2005; Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Oltari, July 26 th, 2005; Apati{an, July 26 th, 2005; Jezera, July 26 th, 2005; [tirova~a, July 27 th, 2005; Polo`ine, July 27 th, 2005; [prenga, July 27 th, 2005; Ljubica, July 27 th, 2005; Sun erac, July 28 th, 2005; Ani}a Luka, August 1 st, 2005; Borisov dom mt. hut, August 2 nd, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; above Borisov dom mt. hut, August 26 th, 2005; Pari}i, August 26 th, 2005; Rami}i, August 26 th, 2005; Bru{ane, July 6 th, 2006; Ba{ke O{tarije, July 7 th, 2006; Bu`im, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; Duplje, July 20 th, 2006; Ba{ke O{tarije, July 21 st, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006; to Sijasetska draga, July 26 th, 2006; forest hut Paklenica NP, July 27 th, 2006; Borisov dom mt. hut, July 27 th, 2006; Jadovno, July 28 th, 2006; Radu~, July 28 th, Argynnis pandora (Denis & Schiffermüller, 1775) (**): turn to Pandore, June 17 th, 2005; [prenga, July 27 th, 2005; Jadovno, July 6 th, 2006.

12 40 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia Argynnis aglaja (Linnaeus, 1758) (* / ** / ***): Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Konjsko, June 18 th, 2005; Babi} Si~a, July 25 th, 2005; Modri} dolac, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Oltari, July 26 th, 2005; Apati{an, July 26 th, 2005; Sun erac, July 28 th, 2005; Trnovac, June 23 rd, 2006; Jadovno, July 6 th, 2006; Bu`im, July 6 th, 2006; Ljubica, July 6 th, 2006; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Petra{ica, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Tudorevo, July 19 th, 2006; Veliki Lubenovac, July 19 th, 2006; Vu~jak, July 20 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006; Malovan (M. stanovi), July 26 th, 2006; Struge, July 27 th, 2006; Jadovno, July 28 th, 2006; Radu~, July 28 th, 2006; Bunovac, July 28 th, Argynnis adippe (Denis & Schiffermüller, 1775) (* / ** / ***): Babi} Si~a, July 25 th, 2005; Apati{an, July 26 th, 2005; Jezera, July 26 th, 2005; [tirova~a, July 27 th, 2005; Dabarska kosa, July 27 th, 2005; Sun erac, July 28 th, 2005; Mala Mo~ila, August 2 nd, 2005; Jadovno, July 6 th, 2006; Bakovac, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Tudorevo, July 19 th, 2006; Veliki Lubenovac, July 19 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; [arinac, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; Jadovno, July 21 st, Issoria lathonia (Linnaeus, 1758) (* / ** / ***): Ljubica, June 18 th, 2005; Konjsko, June 18 th, 2005; [tirova~a, June 18 th, 2005; Borova~ki Pade`, June 18 th, 2005; [vica, July 25 th, 2005; Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Apati{an, July 26 th, 2005; Jezera, July 26 th, 2005; [tirova~a, July 27 th, 2005; Polo`ine, July 27 th, 2005; Borisov dom mt. hut, August 2 nd, 2005; to Bu`im, July 6 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Glamo~eva Duliba, July 20 th, 2006; Bunovac, July 26 th, 2006; Bunovac, July 28 th, Brenthis ino (Rottemburg, 1775) (* / **): Sun erac, July 28 th, 2005; Trnovac, June 20 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, Brenthis daphne (Denis & Schiffermüller, 1775) (* / ** / ***): Babi} Si~a, July 25 th, 2005; Apati{an, July 26 th, 2005; [tirova~a, July 27 th, 2005; [prenga, July 27 th, 2005; Sun erac, July 28 th, 2005; Trnovac, June 23 rd, 2006; Bu`im, July 6 th, 2006; Bru{ane, July 6 th, 2006; Bu`im, July 7 th, 2006; Bakovac, July 7 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; [arinac, July 20 th, 2005; Te`akova~ko vrelo, July 21 st, 2006; Bunovac, July 28 th, Brenthis hecate (Denis & Schiffermüller, 1775) (* / **): Vu~jak, July 25 th, 2005; Modri} dolac, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Apati{an, July 26 th, 2005; Bu`im, July 7 th, Vodice, July 20 th, 2006; [arinac, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006.

13 Nat. Croat. Vol. 16(1), Boloria euphrosyne (Linnaeus, 1758) (* / **): Babi} Si~a, June 17 th, 2005; to Oltari, June 17 th, 2005; Ljubica, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; Sun erac, July 28 th, Vodice, July 20 th, Boloria dia (Linnaeus, 1767) (**): Jadovno, July 21 st, Nymphalinae Nymphalini Vanessa atalanta (Linnaeus, 1758) (* / ** / ***): Ljubica, June 18 th, 2005; Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Jezera, July 26 th, 2005; Sun erac, July 28 th, 2005; Ani}a Luka, August 1 st, 2005; Rami}i, August 26 th, 2005; Kosinjski Bakovac, July 19 th, 2006; Kriva~a, July 20 th, Vodice, July 20 th, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; to Sijasetska draga, July 26 th, 2006; Malovan (M. stanovi), July 26 th, 2006; Jadovno, July 28 th, Vanessa cardui (Linnaeus, 1758) (* / ** / ***): Dabarska kosa, June 18 th, 2005; Borova~ki Pade`, June 18 th, 2005; [tirova~a, July 27 th, 2005; Kubus, July 28 th, 2005; Jadovno, July 6 th, 2006; to Bu`im, July 6 th, 2006; Bu`im, July 6 th, 2006; Bru{ane, July 6 th, 2006; Ljubica, July 6 th, 2006; Ba{ke O{tarije, July 7 th, 2006; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; to Klementa, July 7 th, 2006; Petra{ica, July 7 th, 2006; [tirova~a, July 7 th, 2006; from Begova~a to Krasno, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Tudorevo, July 19 th, 2006; Vu~jak, July 20 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; [arinac, July 20 th, 2006; Ba{ke O{tarije, July 21 st, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; to Sijasetska draga, July 26 th, 2006; Bunovac, July 26 th, 2006; Malovan (M. stanovi), July 26 th, 2006; Malovansko jezero, July 26 th, 2006; forest hut Paklenica NP, July 27 th, 2006; Jadovno, July 28 th, 2006; Bunovac, July 28 th, Inachis io (Linnaeus, 1758) (* / ** / ***): Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Apati{an, July 26 th, 2005; [tirova~a, July 27 th, 2005; [prenga, July 27 th, 2005; Ljubica, July 27 th, 2005; Sun erac, July 28 th, 2005; Mala Mo~ila, August 2 nd, 2005; Ljubica, July 6 th, 2006; Kosinjski Bakovac, July 19 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Tudorevo, July 19 th, 2006; Veliki Lubenovac, July 19 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006; to Sijasetska draga, July 26 th, 2006; Bunovac, July 26 th, 2006; Malovansko jezero, July 26 th, Aglais urticae (Linnaeus, 1758) (* / **): Babi} Si~a, June 17 th, 2005; turn to Pandore, June 17 th, 2005; Vu~jak, June 17 th, 2005; [tirova~a, June 18 th, 2005; Vu~jak, July 25 th, 2005; Apati{an, July 26 th, 2005; to Kugina

14 42 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia ku}a mt. hut, July 28 th, 2005; Tudorevo, July 19 th, 2006; Vu~jak, July 20 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, 2006; Ba{ke O{tarije, July 21 st, Polygonia c-album (Linnaeus, 1758) (* / ** / ***): Dabarska kosa, June 18 th, 2005; Mala Mo~ila, August 2 nd, 2005; Kosinjski Bakovac, July 19 th, 2006; Veliki Lubenovac, July 19 th, Vodice, July 20 th, 2006; Jadovno, July 21 st, 2006; Jadovno, July 28 th, Polygonia egea (Cramer, 1775) (* / ** / ***): turn to Zavi`an mt. hut, July 26 th, 2005; Apati{an, July 26 th, 2005; Sun erac, July 28 th, 2005; Ani}a Luka, August 1 st, 2005; above Borisov dom mt. hut, August 26 th, Vodice, July 20 th, 2006; Jadovno, July 21 st, Araschnia levana (Linnaeus, 1758) (**): Jadovno, July 21 st, Nymphalis antiopa (Linnaeus, 1758) (**): Apati{an, July 26 th, Nymphalis polychloros (Linnaeus, 1758) (**): Ljubica, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; Konjsko, June 18 th, Melitaeini Euphydryas aurinia (Rottemburg, 1775) (**): Ljubica, June 18 th, 2005; Trnovac, June 20 th, Melitaea cinxia (Linnaeus, 1758) (**): [vica, June 16 th, 2005; Ljubica, June 18 th, Melitaea phoebe (Denis & Schiffermüller, 1775) (**): Trnovac, June 23 rd, 2006; Jadovno, July 6 th, 2006; Ljubica, July 6 th, 2006; Jadovno, July 21 st, Melitaea trivia (Denis & Schiffermüller, 1775) (**): to Bakovac, July 7 th, 2006; Bakovac, July 7 th, Melitaea didyma (Esper, 1778) (**): [vica, June 16 th, 2005; turn to Pandore, June 17 th, 2005; Ljubica, June 18 th, 2005; Jadovno, July 6 th, 2006; to Bakovac, July 7 th, Melitaea diamina (Lang, 1789) (**): Trnovac, June 20 th, 2006; Jadovno, July 6 th, Melitaea aurelia Nickerl, 1850 (**): [vica, June 16 th, 2005; Ljubica, June 18 th, 2005; Borova~ki Pade`, June 18 th, 2005; Trnovac, June 23 rd, 2006; Jadovno, June 23 rd, 2006; Jadovno, July 6 th, 2006; Petra{ica, July 7 th, 2006.

15 Nat. Croat. Vol. 16(1), Melitaea britomartis Assmann, 1847: [vica, June 16 th, Melitaea athalia (Rottemburg, 1775) (* / **): turn to Pandore, June 17 th, 2005; Trnovac, June 23 rd, 2006; Jadovno, July 6 th, 2006; to Bu`im, July 6 th, 2006; Bu`im, July 6 th, 2006; Ljubica, July 6 th, 2006; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Tudorevo, July 19 th, 2006; Veliki Lubenovac, July 19 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Ba~i} Duliba, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006; Jadovno, July 28 th, Limenitinae Limenitis populi (Linnaeus, 1758) (**): Kosinjski Bakovac, July 19 th, Limenitis reducta Staudinger, 1901 (* / ** / ***): Gornja Klada, June 17 th, 2005; to Dabarska kosa, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; Oltari, July 26 th, 2005; Ljubica, July 27 th, 2005; Borisov dom mt. hut, August 2 nd, 2005; Mala Mo~ila, August 2 nd, 2005; Rami}i, August 26 th, 2005; Kosinjski Bakovac, July 19 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006; Ani}a Luka, July 27 th, 2006; forest hut Paklenica NP, July 27 th, 2006; Borisov dom mt. hut, July 27 th, Neptis sappho (Pallas, 1771) (**): to Dabarska kosa, June 18 th, Apaturinae Apatura iris (Linnaeus, 1758) (** / ***): Jazmaku{a, July 21 st, 2006; Struge, July 27 th, Satyrinae Elymniini Pararge aegeria (Linnaeus, 1758) (* / ***): Ani}a Luka, August 2 nd, 2005; Ani}a Luka, August 27 th, 2005; Glamo~eva Duliba, July 20 th, 2006; Pari}i, July 27 th, Lasiommata megera (Linnaeus, 1767) (** / ***): [prenga, July 27 th, 2005; to Mala Mo~ila, August 2 nd, 2005; above Borisov dom mt. hut, August 26 th, 2005; Pari}i, August 26 th, 2005; Ba{ke O{tarije, July 21 st, 2006; Dabarska kosa, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, Lasiommata maera (Linnaeus, 1758) (* / **): to Oltari, June 17 th, 2005; to Dabarska kosa, June 18 th, 2005; Polo`ine, July 27 th, 2005; Kubus, July 28 th, 2005; Bu`im, July 7 th, 2006; Bakovac, July 7 th, 2006; from Begova~a to Krasno, July 7 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Tudorevo,

16 44 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia July 19 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, Coenonymphini Coenonympha rhodopensis Elwes, 1900 (* / **): Babi} Si~a, June 17 th, 2005; Dabarska kosa, June 18 th, 2005; Borova~ki Pade`, June 18 th, Coenonympha arcania (Linnaeus, 1761) (* / ** / ***): to Oltari, June 17 th, 2005; turn to Pandore, June 17 th, 2005; Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; Borova~ki Pade`, June 18 th, 2005; Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Apati{an, July 26 th, 2005; [prenga, July 27 th, 2005; Sun erac, July 28 th, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; Trnovac, June 20 th, 2006; Jadovno, June 20 th, 2006; Jadovno, June 23 rd, 2006; ^a~i} Draga, July 6 th, 2006; Bru{ane, July 6 th, 2006; Ba{ke O{tarije, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Cigani{te, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Jadovno, July 21 st, Coenonympha glycerion (Borkhausen, 1788) (* / **): Apati{an, July 26 th, 2005; Sun erac, July 28 th, 2005; Trnovac, June 23 rd, 2006; Jadovno, June 23 rd, 2006; Jadovno, July 6 th, 2006; to Bu`im, July 6 th, 2006; Ljubica, July 6 th, 2006; Ba{ke O{tarije, July 7 th, 2006; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Petra{ica, July 7 th, Coenonympha pamphilus (Linnaeus, 1758) (* / ** / ***): [vica, June 16 th, 2005; Babi} Si~a, June 17 th, 2005; to Oltari, June 17 th, 2005; turn to Pandore, June 17 th, 2005; Gornja Klada, June 17 th, 2005; Ljubica, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; Borova~ki Pade`, June 18 th, 2005; Pari}i, August 26 th, 2005; Trnovac, June 20 th, 2006; Bakovac, July 7 th, 2006; Kri`i}i, July 20 th, 2006; Cigani{te, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Malovan (M. stanovi), July 26 th, Maniolini Aphantopus hyperantus (Linnaeus, 1758) (**): Kosinjski Bakovac, July 19 th, Maniola jurtina (Linnaeus, 1758) (* / ** / ***): [vica, June 16 th, 2005; turn to Pandore, June 17 th, 2005; Dabarska kosa, June 18 th, 2005; [vica, July 25 th, 2005; Babi} Si~a, July 25 th, 2005; turn to Pandore, July 26 th, 2005; Dabarska kosa, July 27 th, 2005; Kubus, July 28 th, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; above Borisov dom mt. hut, August 26 th, 2005; Pari}i, August 26 th, 2005; Trnovac, June 20 th, 2006; Jadovno, June 23 rd, 2006; Bu`im, July 6 th, 2006; ^a~i} Draga, July 6 th, 2006; Ljubica, July 6 th, 2006; Ba{ke O{tarije, July 7 th, 2006; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Ba~i} Duliba, July 21 st, 2006; Jadovno, July 21 st, 2006; Jadovno, July 21 st, 2006; Veliki Javornik, July 27 th, 2006.

17 Nat. Croat. Vol. 16(1), Hyponephele lycaon (Rottemburg, 1775) (*): [arinac, July 20 th, 2006; Babrova~a, July 20 th, Hyponephele lupinus (O. Costa, 1836) (**): Oltari, July 26 th, Erebiini Erebia ligea (Linnaeus, 1758) (* / ** / ***): Babi} Si~a, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Apati{an, July 26 th, 2005; Polo`ine, July 27 th, 2005; Sun erac, July 28 th, 2005; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; to Klementa, July 7 th, 2006; from Begova~a to Krasno, July 7 th, 2006; above Dundovi} Pade`, July 19 th, 2006; Veliki Lubenovac, July 29 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; Duplje, July 20 th, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Malovan (M. stanovi), July 26 th, 2006; Bunovac, July 28 th, Erebia euryale (Esper, 1805) (* / **): Babi} Si~a, July 25 th, 2005; Tudorevo, July 19 th, 2006; Ba~i} Duliba, July 21 st, 2006; Jadovno, July 21 st, 2006; Jadovno, July 28 th, Erebia epiphron (Knoch, 1783) (* / **): Modri} dolac, July 25 th, 2005; Jezera, July 26 th, 2005; Tudorevo, July 19 th, 2006; Kri`i}i, July 20 th, Erebia aethiops (Esper, 1777) (** / ***): Polo`ine, July 27 th, 2005; [prenga, July 27 th, 2005; Velika Mo~ila, August 2 nd, 2005; after Bru{ane, July 6 th, Erebia medusa (Denis & Schiffermüller, 1775) (* / ** / ***): [vica, June 16 th, 2005; Babi} Si~a, June 17 th, 2005; to Oltari, June 17 th, 2005; turn to Pandore, June 17 th, 2005; Ljubica, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; Borova~ki Pade`, June 18 th, 2005; Ljubica, July 6 th, 2006; Petra{ica, July 7 th, 2006; [tirova~a, July 7 th, 2006; Tudorevo, July 19 th, 2006; Kriva~a, July 20 th, 2006; Malovan (M. stanovi), July 26 th, Erebia ottomana Herrich-Schäffer, 1847 (***): Bunovac, July 26 th, 2006; to Malovan (M. stanovi), July 26 th, 2006; Malovan (M. stanovi), July 26 th, 2006; Malovansko jezero, July 26 th, 2006; Struge, July 27 th, 2006; Bunovac, July 28 th, Erebia melas (Herbst, 1796) (* / ***): above Dundovi} Pade`, July 19 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Malovan (M. stanovi), July 26 th, 2006; Malovansko jezero, July 26 th, 2006; Babino jezero, July 28 th, Erebia oeme (Hübner, 1804) (* / ** / ***): Jezera, July 26 th, 2005; [tirova~a, July 27 th, 2005; Sun erac, July 28 th, 2005; Ljubica,

18 46 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia July 6 th, 2006; Petra{ica, July 7 th, 2006; [tirova~a, July 7 th, 2006; Malovan (M. stanovi), July 26 th, Melanargiini Melanargia galathea (Linnaeus, 1758) (* / ** / ***): to Dabarska kosa, June 18 th, 2005; [vica, July 25 th, 2005; Babi} Si~a, July 25 th, 2005; Modri} dolac, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Oltari, July 26 th, 2005; turn to Pandore, July 26 th, 2005; Jezera, July 26 th, 2005; Apati{an, July 26 th, 2005; [prenga, July 27 th, 2005; Dabarska kosa, July 27 th, 2005; Ljubica, July 27 th, 2005; Sun erac, July 28 th, 2005; Kubus, July 28 th, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; Jadovno, July 6 th, 2006; to Bu`im, July 6 th, 2006; Bu`im, July 6 th, 2006; Ljubica, July 6 th, 2006; Bu`im, July 7 th, 2006; to Bakovac, July 7 th, 2006; Bakovac, July 7 th, 2006; Petra{ica, July 7 th, 2006; Krasno, July 7 th, 2006; Kosinjski Bakovac, July 19 th, 2006; Kri`i}i, July 20 th, 2006; Kriva~a, July 20 th, 2006; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Cigani{te, July 20 th, 2006; [arinac, July 20 th, 2006; Babrova~a, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; to Kugina ku}a mt. hut, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006; Bunovac, July 26 th, 2006; Jadovno, July 28 th, 2006; Bunovac, July 28 th, Satyrini Satyrus ferula (Fabricius, 1793) (*): Tudorevo, July 19 th, 2006; Glamo~eva Duliba, July 20 th, Hipparchia fagi (Scopoli, 1763) (* / ** / ***): Sun erac, July 28 th, 2005; Borisov dom mt. hut, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; Glamo~eva Duliba, July 20 th, Vodice, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006; Pari}i, July 27 th, Hipparchia syriaca (Staudinger, 1871) (* / ** / ***): [vica, July 25 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Oltari, July 26 th, 2005; to Mala Mo~ila, August 2 nd, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; Pari}i, August 26 th, 2005; Rami}i, August 26 th, 2005; to Bakovac, July 7 th, 2006; Babrova~a, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Borisov dom mt. hut, July 27 th, Hipparchia semele (Linnaeus, 1758) (* / ** / ***): to Oltari, June 17 th, 2005; Ljubica, June 18 th, 2005; to Dabarska kosa, June 18 th, 2005; Dabarska kosa, June 18 th, 2005; [tirova~a, June 18 th, 2005; Borova~ki Pade`, June 18 th, 2005; turn to Zavi`an mt. hut, July 26 th, 2005; Oltari, July 26 th, 2005; [prenga, July 27 th, 2005; Dabarska kosa, July 27 th, 2005; Sun erac, July 28 th, 2005; Ani}a Luka, August 1 st, 2005; Mala Mo~ila, August 2 nd, 2005; Velika Mo~ila, August 2 nd, 2005; Pari}i, August 26 th, 2005; Rami}i, August 26 th, 2005; Jadovno, July 6 th, Vodice, July 20 th, 2006; Babrova~a, July 20 th, 2006; Duplje, July 20 th, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; Te`akova~ko vrelo, July 21 st, 2006; Jadovno, July 21 st, 2006.

19 Nat. Croat. Vol. 16(1), Arethusana arethusa (Denis & Schiffermüller, 1775) (***): Pari}i, August 2 nd, 2005; Rami}i, August 26 th, Brintesia circe (Fabricius, 1775) (* / ** / ***): Babi} Si~a, July 25 th, 2005; Oltari, July 26 th, 2005; turn to Pandore, July 26 th, 2005; Dabarska kosa, July 27 th, Vodice, July 20 th, 2006; Ba{ke O{tarije, July 21 st, 2006; Dabarska kosa, July 21 st, 2006; Ba~i} Duliba, July 21 st, 2006; Struge, July 27 th, 2006; Jadovno, July 28 th, Chazara briseis (Linnaeus, 1764) (* / **): Dabarska kosa, July 27 th, Vodice, July 20 th, 2006; Babrova~a, July 20 th, Biogeographical characteristics of species determined on Mt. Velebit are given in Table 2 and Table 3. Biogeographical anaysis has revealed the dominance of the Eurosibirean (62.26%), Oriental (14.15%) and Mediterranean (13.21%) biogeographical elements. DISCUSSION Faunistic analysis The total of 106 butterfly species found on Mt. Velebit in 2005 and 2006 represents 55.80% of the Croatian butterfly fauna, which has altogether 190 species (MIHOCI et al., 2005, 2006; PERKOVI], 2006) or 72.10% of all detected species in the Velebit area until now (Appendix 1, KU^INI] et al., 1995; JUTZELER et al., 2001, 2002). According to the KU^INI] et al. (1995) only Zerynthia polyxena from the Papilionidae family found in Krasno in the past century was not detected in our research, probably because our first fieldtrip was done in June and the species had already emerged. From the Pieridae family we have not detected Pontia daplidice and Colias hyale. Specimens of the species Leptidea morsei ssp. major from the Central Butterfly Collection (CNHM) mentioned in KU^INI] et al. (1995) was misidentified with the species Leptidea sinapis, as confirmed by analysis of the following genitalia parameters: length of the aedeagus, saccus, uncus and tegumen (LELO, 2003) (redet. M. Ku~ini}). According to LELO (2003), data on the metrical values of tegument are best in proving the existence of the explicit differences in genitalia of the L. sinapis and L. morsei ssp. major taxa. It is supposed that the main limiting factor for distribution of L. morsei in Europe is a cooler and rainier summer climate (LORKOVI], 1975). Also, L. morsei inhabits oak forests with the food plants spring vetch Lathyrus verna or black pea L. niger (LORKOVI], 1975). Although both plants are known from Mt. Velebit (FORENBACHER, 2001), verna from Vratnik, O{tarije, Meda~ka staza, Malovan, Debeli kuk and Crnopac and niger from Vratnik, Budakovo brdo, Kiza and Takalice they were not noticed during our fieldtrips.

20 48 Mihoci, I. et al.: Contribution to the butterfly fauna of the Velebit Mountain, Croatia Tab. 2. Biogeographical characteristics of species identified on Mt. Velebit, according to HRUBY (1964), SIJARI] et al. (1984) and HIGGINS & RILEY (1993). Ord. No. Species and author Biogeog. character. 1. Erynnis tages (Linnaeus, 1758) ES 2. Carcharodus floccifera (Zeller, 1847) ME 3. Thymelicus lineola (Ochsenheimer, 1808) ES 4. Thymelicus sylvestris (Poda, 1761) ES 5. Hesperia comma (Linnaeus, 1758) ES 6. Ochlodes venata (Bremer & Grey, 1853) ES 7. Parnassius mnemosyne (Linnaeus, 1758) OR 8. Parnassius apollo (Linnaeus, 1758) ES 9. Iphiclides podalirius (Linnaeus, 1758) ES 10. Papilio machaon Linnaeus, 1758 ES 11. Leptidea sinapis (Linnaeus, 1758) ES 12. Antocharis cardamines (Linnaeus, 1758) ES 13. Aporia crataegi (Linnaeus, 1758) ES 14. Pieris brassicae (Linnaeus, 1758) ES 15. Pieris mannii (Mayer, 1851) OR 16. Pieris rapae (Linnaeus, 1758) ES 17. Pieris ergane (Geyer, 1828) OR 18. Pieris napi (Linnaeus, 1758) HA 19. Colias croceus (Fourcroy, 1785) TR 20. Gonepteryx rhamni (Linnaeus, 1758) ES 21. Gonepteryx cleopatra (Linnaeus, 1767) ME 22. Hamearis lucina (Linnaeus, 1758) ME 23. Lycaena virgaureae (Linnaeus, 1758) ES 24. Lycaena alciphron (Rottemburg, 1775) ES 25. Lycaena candens (Herrich-Schäffer, 1844) OR 26. Satyrium w-album (Knoch, 1782) ME 27. Satyrium pruni (Linnaeus, 1758) ME 28. Satyrium spini (Denis & Schiffermüller, 1775) ES 29. Satyrium ilicis (Esper, 1779) ME 30. Cupido minimus (Fuessly, 1775) ES 31. Cupido alcetas (Hoffmannsegg, 1804) ES 32. Celastrina argiolus (Linnaeus, 1758) ES 33. Scolitantides orion (Pallas, 1771) ES 34. Glaucopsyche alexis (Poda, 1761) ES 35. Maculinea arion (Linnaeus, 1758) ES