FAMILY CANTHARIDAE (Soldier beetles)

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1 FAMILY CANTHARIDAE (Soldier beetles) J. McNamara This family includes 126 species in Canada and Alaska. The adults are common on goldenrod, milkweed, and hydrangea where they feed on nectar and pollen. Most larvae are carnivorous generally preying on other soft-bodied insects; a few are omnivorous, feeding on plants such as wheat, potatoes, celery, and other plants. They occur mostly beneath bark and debris. Some of the genera of Cantharidae are poorly known taxonomically, including the large genus Podabrus, and are in need of revision. Fender (1962) provided keys to the species of the Pacific Northwest. AK (15); YK (8); NT (13); BC (57); AB (20); SK (9); MB (22); ON (48); PQ (45); NB (22); NS (25); PE (1); LB (5); NF (15); I (2) Subfamily CANTHARINAE Tribe Podabrini Genus PODABRUS Westwood Brachynotus Kirby Malthacus Kirby Key to North American species: Fall (1927) P. alexanderi Fender AB P. brevicollis Fall ON P. cavicollis cavicollis LeConte BC P. conspiratus Fall AK - - BC P. deceptus W.J.Brown AK YK NT BC - - MB P. diadema (Fabricius) ON PQ NB NS - - NF P. excursus excursus Fall BC AB P. extremus LeConte AK YK NT BC AB - MB ON LB - P. falli Hopping BC AB P. fissilis Fall AK - - BC AB P. flavicollis LeConte ON P. frater LeConte ON quadricollis (Motschulsky) P. frosti Fender ON - NB P. fumiganus Green PQ NB P. furtivus Fall BC P. heteronychus Fall AK YK NT BC AB SK MB ON PQ P. instabilis Fall BC P. intrusus Green ON PQ NB NS - - NF P. laevicollis (Kirby) AK YK NT BC AB SK MB ON PQ NB - - LB NF P. limbellus LeConte PQ NB P. lutosus LeConte BC P. lygarius Fender BC P. melvillei Fender BC P. modestus (Say) ON PQ - NS P. nothoides LeConte MB ON PQ P. obscurevittatus Fall - YK NT BC AB SK MB P. pattoni LeConte PQ NB P. perplexus W.J.Brown AK - NT MB P. piniphilus (Eschscholtz) AK - NT BC AB - MB - PQ P. planulus Green MB ON PQ - NS - - NF P. probus Fall PQ P. protensus LeConte ON P. pruinosus atrocervicus Fender BC

2 P. pruinosus diversipes Fall BC AB P. puberulus LeConte MB ON PQ NB NS - LB NF P. punctatus LeConte PQ NB NS P. puncticollis (Kirby) PQ NB marginellus LeConte P. punctulatus LeConte ON PQ P. pustulatus Fender BC P. pygmaeus Green PQ P. rossi Fender BC P. rugosulus LeConte ON PQ NB NS parvicollis (Motschulsky) P. scaber LeConte BC AB P. secretus W.J.Brown PQ P. sericatus (Mannerheim) AK P. simplex Couper MB ON PQ NB NS P. smithi Fender BC P. tetragonoderus Fall AK - - BC AB P. tomentosus (Say) ON cinereipennis Motschulsky rufiolus (Melsheimer) P. tricostatus (Say) ON PQ atripes (Motschulsky) benneti (Kirby) Tribe Cantharini Genus ANCISTRONYCHA Märkel Key to North American species: McKey-Fender (1950) A. bilineata (Say) ON PQ - NS A. dentigera (LeConte) ON PQ A. neglecta (Fall) ON PQ Genus CANTHARIS Linné Cyrtomoptila Motschulsky Telephorus Schaeffer C. alticola (LeConte) BC C. aneba borealis McKey-Fender - - NT BC AB SK MB C. beta Fender BC C. consors (LeConte) BC tibialis (LeConte) tibiella (Gemminger) C. curtisi (Kirby) - YK NT BC AB SK MB ON PQ NB - - LB NF samouellei (Kirby) C. impressa (LeConte) ON PQ - NS C. oregona (LeConte) BC C. rotundicollis Say ON PQ NB NS C. rufa Linné ON PQ NB NS PE - NF C. transmarina transmarina (Motschulsky) BC C. transmarina scopa (LeConte) BC C. tuberculata (LeConte) PQ armigera (Couper) brevicollis (LeConte)

3 C. westwoodi (Kirby) Genus CYRTOMOPTERA Motschulsky C. divisa (LeConte) BC latiuscula Motschulsky Genus RHAXONYCHA Motschulsky Key to North American species: McKey-Fender (1950) R. bilobata (McKey-Fender) MB ON PQ R. carolina (Fabricius) ON PQ jactata (Say) Genus RHAGONYCHA Eschscholtz Key to North American species: Green (1940) R. alaskensis Fender AK R. anthracina Mannerheim AK R. costipennis (LeConte) ON R. excavata (LeConte) BC - - MB ON PQ - NS R. fraxini (Say) AK - - BC AB - MB ON PQ - NS - - NF atra (Kirby) binodula Mannerheim nigrita (LeConte) R. fulva (Scopoli) BC R. gilvipes (Gemminger) - - NT -? dichroa (LeConte) flavipes (LeConte) R. imbecillis (LeConte) ON NF albolineata (Blatchley) R. luteicollis (Germar) ON cinctella (LeConte) R. mandibularis (Kirby) AK YK NT BC AB SK MB - PQ LB NF nigritula (LeConte) R. mollis mollis Fall SK - ON - - NS - - NF nigriceps (LeConte) pusilla (LeConte) R. nanula (LeConte) NS R. oriflava (LeConte) ON R. recta (Melsheimer) BC AB - - ON PQ - NS - - NF pusilla (LeConte) rufipes (Say) R. scitula (Say) PQ - NS R. septentrionis (Green) MB ON R. sylvatica (Green) ON R. tantilla (LeConte) SK pusio (LeConte) R. vilis (LeConte) ON PQ R. walshi (LeConte) MB Tribe Silini

4 Genus POLEMIUS LeConte P. canadensis W.J.Brown NB NS P. laticornis (Say) ON dubius (Melsheimer) incisus LeConte invalidus (Say) undulatus (LeConte) Genus SILIS Charpentier Key to North American species: Green (1966) (subgenus SILIS s.str.) S. atra LeConte - - NT S. difficilis difficilis LeConte - YK NT BC AB SK MB - PQ NB S. insperata Green BC S. lecontei Green BC S. lutea LeConte BC pallens LeConte S. pallida Mannerheim AK - - BC S. parallela Green BC S. percomis (Say) ON PQ NB NS curta (LeConte) longicornis LeConte S. spinigera LeConte BC minuta LeConte S. striatella Green BC (subgenus DITEMNUS LeConte) S. bidentata (Say) PQ S. latiloba Blatchley MB Tribe Malthinini Genus MALTHINUS Latreille Key to North American species: Fender (1972) M. occipitalis LeConte ON PQ Genus MALTHODES Kiesenwetter Key to North American species: Fender (1951) M. abruptus Fender BC M. acuticauda Fender BC M. alexanderi Fender BC M. bakeri Fender BC AB M. canadensis Fender BC M. columbiensis Fender BC M. fragilis (LeConte) ON PQ NB NS transversus (LeConte)

5 M. fuliginosus fuliginosus LeConte PQ NB NS M. fuliginosus flavicollis Fender NS M. intricatus Fender BC AB M. macnabi Fender BC M. medioccidens Fender ON M. niger (LeConte) PQ NB NF M. oregonus Fender BC M. parvulus (LeConte) ON PQ NB NS - - NF M. piceolus Fall BC M. piperi Fender BC M. similis Fender ON - - NS M. stacesmithi Fender BC M. terraenovae Fender NF Subfamily CHAULIOGNATHINAE Tribe Chauliognathini Genus CHAULIOGNATHUS Hentz Key to North American species: Fender (1964) C. pennsylvanicus (DeGeer) MB ON bimaculatus (Fabricius) Tribe Ichthyurini Genus TRYPHERUS LeConte Lygerus Kiesenwetter Key to North American species: Fender (1960) T. frisoni Fender ON

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