Checklist of Diving beetles (Insecta, Coleoptera, Dytiscidae) of Georgia
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1 Proceedings of the Institute of Zoology, 2008, vol. 23. P Checklist of Diving beetles (Insecta, Coleoptera, Dytiscidae) of Georgia G. Chaladze Institute of zoology, Georgia, Abstract. Last checklist of Diving Beetles of Georgia was published in 1953 by F. Zaitzev and included 80 species; since then taxonomic changes occurred and some new species were registered and described. For this reason, it was necessary to update records. Currently there are 25 genera and 85 species registered in Georgia. Abbreviations: EG - East Georgia; WG West Georgia. Key words: Diving beetles, Coleoptera, Distribution, Georgia. 1. L. haemorrhoidalus (Fabricius, 1787) Subfamily Copelatinae Branden, 1885 Genus Liopterus Dejean, 1833 Subfamily Laccophilinae Gistel, 1856 Genus Laccophilus Leach in Brewster, L. hyalinus (De Geer, 1774) 3. L. minutus (Linnaeus, 1758) 4. L. poecilus Klug, 1834 Distribution: EG: Tbilisi, Baraleti, Dzveli Lake, Manglisi WG: Anaklia[4, 7, and Personal data] Subfamily Hydroporinae Aube, 1836 Genus Hyphydrus Illiger, H. ovatus (Linnaeus, 1761) Distribution: EG: Gvileti, Kodjori, Mukhrovani [3,4]. 6. H. cuspidatus (Kunze, 1818) Distribution: EG:Gardabani [3,4]. Genus Hydrovatus Motschulsky, 1853 Genus Hydroglyphus Motschulsky, H. geminus (Fabricius, 1792) Distribution: All over Gerogia [3,4]. 8. H. signatellus (Klug, 1834) Distribution: EG: Tbilisi, Mukhrovani, Gardabani, Akhaltsikhe, Mataniskhevi [4].
2 Genus Hygrotus Stephens, H. armeniacus (Zaitzev, 1927) Distribution: EG: Baraleti [4]. 10. H. confluens (Fabricius, 1787) Distribution: EG: Tbilisi, Gardabani, Mukhrovani, Manglisi, Borjomi [4]. 11. H. enneagrammus (Ahrens, 1833) Distribution: EG: Tbilisi, Gardabani, Mtskheta, Mukhrovani, Manglisi [4]. 12. H. flaviventris (Motschulsky, 1860) Distribution: Eg: Tbilisi, Gardabani, Mukhrovani [4]. 13. H. impressopunctatus (Schaller, 1783) 14. H. lernaeus Schaum, 1857 Distribution: EG: Lake Kus Tba [3]. 15. H. marklini (Gyllenhal, 1813) Distribution: EG: Bakuriani, Baraleti, Tabatskuri Lake [4]. 16. H. pallidus (Aube, 1850) Distribution: EG: Tbilisi [4,7]. 17. H. inaequalis (Fabricius, 1776) Genus Herophydrus Sharp, H. musicus (Klug, 1834) Distribution: EG: Tbilisi, Gardabani, Mukhrovani [4]. Genus Deronectes Sharp, D. parvicollis Schaum, 1864 Distribution: EG:Tbilisi, Kojori, Bakuriani, Manglisi, Gardabani [4]. Genus Nebrioporus Regimbart, N. airumlus (Kolenati, 1845) Distribution: EG: Bakuriani, Borjomi, Mleta, Pasanauri, Gvileti, Gori, Manglisi, Tbilisi, Gardabani [4]. 21. N. suavis (Sharp, 1882) Distribution: EG: Tbilisi [4]. Genus Stictotarsus Zimmermann, S. griseostriatus (DeGeer, 1774) Distribution: Bakuriani, Tsemi, Tabatskuri Lake [4]. Genus Scarodytes Gozis, S. halensis halensis (Fabricius, 1787) Distribution: EG: Bakuriani, Baraleti, Ktsia, Tsemi, Tabatskuri Lake [4]. 24. O. davisii davisii (Curtis, 1831) Distribution: EG: Mleta [4]. Genus Oreodytes Seidlitz, 1887 Genus Graptodytes Seidlitz, 1887
3 25. G. bilineatus Sturm, 1835 Distribution: WG: Batumi [4]. Genus Porhydrus Guignot, P. lineatus (Fabricius, 1775) Distribution: EG: Tbilisi, Mukhrovani [4]. Genus Hydroporus Clairville, H. erythrocephalus (Linnaeus, 1758) Distribution: EG: Tsalka, Bakuriani, Tabatskuri Lake [4]. 28. H. angustatus Sturm, 1835 Distribution: EG: Tsalka [4]. 29. H. palustris (Linnaeus, 1761) Distribution: EG: Bakuriani, Borjomi, Tabatskuri Lake, Tbadzveli, Tsalka, Manglisi [4]. 30. H. jonicus caucasicus Zaitzev, 1927 Distribution: EG: Gvileti [4]. 31. H. ampliatus colchicus Bilyashiwsky, H. nigellus Mannerheim, 1853 Distribution: EG: Ktsia, Bakuriani, Baraleti, Tabatskuri Lake [4]. 33. H. marganatus Duftschmid, 1805 Distribution: EG: Baraleti, Ktsia [4]. 34. H. planus (Fabricius, 1781) Distribution: EG: Bakuriani, Baraleti, Tbadzveli, Borjomi, Tbilisi, Gardabani [4]. 35. H. tessellatus (Drapiez, 1819) Distribution: EG: Tbilisi, Saguramo, Lagodekhi, Telavi, Akhaltsikhe, WG: Sukhumi, [4]. 36. H. kozlovskii Schaum, 1868 Distribution: EG: Baraleti, Ktsia, Tabatskuri Lake [4]. 37. H. discretus Fairmare & Brisout, 1859 Distribution: EG: Baraleti, Ktsia, Surami, gvileti, Mleta, Pasanauri, Lagodekhi [4]. 38. H. georgicus Bilyashiwsky, H. jacobsoni Zaitzev, 1927 Distribution: EG: Bitschuri, Baraleti, Bakuriani, Gvileti, Lagodekhi [2,4]. 40. H. lencoranensis Fery, H. pubescens (Gyllenhal, 1808) 42. H. thracicus Gueorguiev, 1966 Distribution: EG: Bakuriani [2]. 43. H. transgrediens Gschwendtner, P. maculatus (Linnaeus, 1758) Distribution: EG: Mtskheta, Tbilisi [4]. 45. P. lunulatus (Steven, 1829) Subfamily Agabinae Thomson, 1867 Genus Platambus Thomson, 1859
4 Distribution: EG: Bakuriani, Borjomi, Abastumani, Pasanauri, Manglisi, Lagodekhi, Mleta, Tbilisi [4]. Genus Agabus Leach, A. Caraboides Sharp, 1882 Distribution: EG: Baraleti [4]. 47. A. glacialis Hochhuth, 1846 Distribution: EG: Khevsureti, Manglisi, Bakuriani, Kodjori, Gvileti, GW: Svaneti [4 and Personal data]. 48. A. biguttatus (Olivier, 1795) Syn.: Gaurodytes nitidus Fabricius, 1801 Distribution: EG: Tbilisi, Qvabiskhevi [7 and Personal data]. 49. A. bipustulatus (Linnaeus, 1767) Syn.: Gaurodytes solieri Aube, 1837 Distribution: EG: Tbilisi [4,7]. 50. A. congener (Thunberg, 1794) Distribution: EG: Bakuriani, Baraleti, Borjomi, Tbadzveli, Ktsia, WG: Gagra [4]. 51. A. bergi Zaitzev, 1913 Distribution: EG: Levan-gel Lake [4]. 52. A. amoenus amoenus Solsky, 1874 Distribution: EG: Tbilisi [4]. 53. A. nebulosus (Foster, 1771) Distribution: EG: Tbilisi, Kobi, Manglisi, Gvileti, Telavi, Mukhrovani, Tabatskuri Lake [4,7]. 54. A. conspersus (Marsham, 1802) Distribution: EG: Tbilisi, Bazaleti Lake, Mukhrovani, Gardabani [4,7]. 55. A. sturmii (Gyllenhal in Schonherr, 1808) Distribution: EG: Bakuriani, Tabatskuri Lake [4]. 56. A. coxalis schmidti Zaitzev, 1913 Distribution: EG: Tabatskuri Lake [4]. 57. A. labiatus (Brahm, 1791) Distribution: EG: Tuman-gel Lake [4,7]. 58. A. undulatus (Schrank, 1776) 59. A. affinis (Paykull, 1798) Distribution: EG: Tbilisi [7]. 60. A. dilatatus (Brulle, 1832) 61. A. paludosus (Fabricius, 1801) Distribution: EG: Tsitsamuri [Personal data]. Genus Ilybius Erichson, I. cinctus Sharp, 1882 Distribution: EG: Lake Kus Tba [4]. 63. I. fuliginosus (Fabricius, 1792) Distribution: EG: Baraleti, Borjomi, Manglisi, Gvileti, Tabatskuri Lake, Khevsha [4]. 64. I. satunini (Zaitzev, 1913) Distribution: EG: Bakuriani, Baraleti, Tbadzveli, Ktsia [4]. 65. I. subaeneus Erichson, 1837 Distribution: EG: Tavfaravani Lake, Borjomi, Kobi, Mleta, WG: Gagra [4]. Subfamily Colymbetinae Erichson, 1837
5 Genus Rhantus Dejean, Rh. frontalis (Marsham, 1802) Syn.: Rhantus suturalis Lacordaire, 1835 Distribution: EG: Tsalka, Borjomi, Baraleti, Manglisi, Mukhrovani [4]. 67. Rh. notaticollis (Aube, 1837) Distribution: WG: Poti [4,6]. 68. Rh. suturalis (MacLeay, 1825) Syn.: punctatus Geoffroy, 1785 Distribution: EG: Tbilisi, Mukhrovani, Mtskheta, Signagi, Gardabani, Pasanauri, WG: Poti [4]. Genus Colymbetes Clairville, C. koenigi Zaitzev, 1927 Distribution: EG: Baraleti, Tabatskhuri Lake [4]. 70. C. semenowi (Jakovlev, 1896) Distribution: Georgia [1,5]. Subfamily Dytiscinae Leach in Brewster, 1815 Genus Eretes Laporte, E. griseus Fabricius, 1781 Distribution: EG: Baraleti, Tabatskhuri Lake [4]. Genus Hydaticus Leach, H. transversalis transversalis Pontoppidan, 1763 Distribution: WG: Poti, Gagra, Gudauta, Batumi [4]. 73. H. schelkovnikovi Zaitzev, 1913 Distribution: EG: Gardabani, WG: Poti, Gagra, Sukhumi, Batumi [4]. 74. H. grammicus Germar, 1830 Distribution: EG: Tbilisi, Gardabani, WG: Poti [4,6]. Genus Graphoderus Dejean, G. cinereus Linne, 1758 Distribution: EG: Tbilisi, Manglisi, Bakuriani [4]. 76. G. zonatus Hoppe, 1795 Distribution: EG: Tabatskhuri Lake, Borjomi, Manglisi [4]. Genus Acilius Leach, A. sulcatus (Linnaeus, 1758) Distribution: EG: From Bakuriani to Tbilisi, Gvileti, Gudauri [4]. 78. D. dimidiatus Bergstrasser, 1778 Distribution: WG: Pichori [6]. 79. D. semisulcatus O.F.Muller, 1776 Genus Dytiscus Linnaeus, 1758
6 Distribution: WG: Pichori, Batumi [4]. 80. D. lapponicus lapponicus Gyllenhal, 1808 Distribution: EG: Tabatskhuri Lake [4]. 81. D. persicus Wehncke, 1876 Distribution: Georgia [4]. 82. D. circumcinctus Ahrens, 1811 Distribution: EG: Chartali [Personal data]. 83. D. circumflexus Fabricius, 1801 Distribution: EG: Tabatskhuri Lake [4]. 84. D. lateralimarginalis torquantus Fischer von Waldheim, 1829 Distribution: WG: Poti, Anaklia [4 and Personal data]. 85. D. marginalis marginalis Linnaeus, 1758 Bibliography 1. Nilssons N.A Catalogue of Palaearctic Dytiscidae (web-version 2. Shaverdo, H. V Revision of the nigrita-group of Hydroporus Clairville, 1806 (Insecta: Coleoptera: Dytiscidae). Annalen des Naturhistorischen Museums in Wien Serie B Botanik und Zoologie 105 B: Зайцев Ф.А Плавунцы Кавказа, раб.сев-кав.гидробиол.стан. 2 с Зайцев Ф. А Водные жестокрылие в фауне Грузии. Тр. Зоолю Инст. АН ГССР XI Зайцев Ф.А Плавунцовые и вертячки, Фауна СССР. Жесткокрылые, т. 4. М.-Л., 377 с. 6. Кобахидзе Д. Н Анализ наземных биоценозов центральной части Колхидской низменности. Тр. Зоолю Инст. АН ГССР V., Якобсон Г.Г Жуки России и Западной Европы.. С.-Петербург. с
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