FAUNA OF THE CAVES OF YUCATAN

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FAUNA OF THE CAVES OF YUCATAN By A. S. PEARSE Duke University WITH THE COLLABORATION OF NATHAN BANKS JOSEPH C. BEQUAERT JOSEPH C. CHAMBERLIN RALPH V. CHAMBERLIN B. G. CHITWOOD WILLIAM J. CLENCH EDWIN P. CREASER NORMA C. FURTOS HELEN T. GAIGE THEODORE H. HUBBELL CARL L. HUBBS LIBBIE H. HYMAN WILTON IVIE REMINGTON KELLOGG HARLOW B. MILLS J. PERCY MOORE C. F. W. MUESEBECK et al. GRACE E. PICKFORD HORACE W. STUNKARD GEORGE W. WHARTON WILLIAM MORTON WHEELER CHARLES BRANCH WILSON FREDERICK A. WOLF PUBLISHED EY CARNEGIE INSTITUTION OF WASHINGTON

CARNEGIE INSTITUTION OF WASHINGTON PUBLICATION NO. 491 [This book first issued June 15, 1938] LORD BALTIMORE PRESS, BALTIMORE, MO. STANDARD ENGRAVING CO., WASHINGTON, D. C. MERIDEN GRAVURE CO., MERIDEN, CONN.

AS CONTENTS PAGE Introduction. A. S. PEARSE (8 figs.).......................................... 1 Conditions of life in caves............................................ 1 Caves of Yucatan............................................ 9 1. Fungal Flora of Yucatan Caves. FREDERICK A. WOLF (1 pl.).......... 19 II. Land Planarians from Yucatan. LIBBIE H. HYMAN (9 figs.)....................... 23 III. Parasitic Flatworms from Yucatan. HORACE W. STUNKARD (8 figs.)........... 33 IV. Sorne Nematodes from the Caves of Yucatan. B. G. CHITWOOD (45 figs.)... 51 V. Leeches (Hirudinea) from Yucatan Caves. J. PERCY MOORE (2 figs.)........ 67 VI. Earthworms in Yucatan Caves. GRACE E. PICKFORD (16 figs., 3 pis.)... 71 VII. Arachnida of the Orders of Pedipalpida, Scorpionida, and Ricinulida. RALPH V. CHAMBERLIN and WILTON IVIE (17 figs.)... 101 VIII. A New Genus and Three New Species of False Scorpions from Yucatan Caves (Arachnida-Chelonethida). JOSEPH C. CHAMBERLIN (4 figs.)...... 109 IX. Araneida from Yucatan. RALPH V. CHAMBERLIN and WILTON IVIE (24 figs.)... 123 X. Acarina of Yucatan Caves. G. W. WHARTON (28 figs.)...... 137 XI. Copepoda from Yucatan Caves. A. S. PEARSE with C. B. WILSON................ 153 XII. A New Species of Cypridopsis from Yucatan. NORMA C. FURTOS (1 fig.)... 155 XIII. Large Cave Crustacea of the Yucatan Peninsula. EDWIN P. CREASER (8 figs.).... 159 XIV. Diplopoda from Yucatan. RALPH V. CHAMBERLIN (55 figs.)... 165 XV. Collembola from Yucatan Caves. HARLOW B. MILLS (27 figs.)...... 183 XVI. New Cave-Crickets from Yucatan, with a Review of the Pentacentrinae, and Studies on the Genus Amphiacusta (Orthoptera, Gryllidae). THEODORE H. HUBBELL (78 figs., 2 graphs)... 191 XVII. A New Myrmeleonid from Yucatan. NATHAN BANKS... 235 XVIII. Insects from Yucatan Caves. A. S. PEARSE with C. F. W. MUESEBECK et 0,1.... 237 XIX. Ants from the Caves of Yucatan. WILLIAM MORTON WHEELER.......... 251 XX. A Third Contribution to the Molluscan Fauna of Yucatan. J. BEQUAERT and W. J. CLENCH..................... 257 XXI. Fishes from the Caves of Yucatan. CARL 1. HUBBS (4 pis.)..... 261 XXII. Sorne Reptilian Records from Caves of Yucatan. HELEN T. GAIGE... 297 XXIII. Birds in Yucatan Caves. A. S. PEARSE... 299 XXIV. Mammalia from Yucatan Caves. A. S. PEARSE with REMINGTON KELLOGG... 301 129972 iii

XVII A NEW MYRMELEONID FROM YUCATAN NATHAN BANKS Specimens of an ant-lion fly and the larvae collected in caves in Yucatan occurred near the entrance of the caves. These specimens prove to belong to the genus Eremoleon, one species of which was known from Cuba, and one from Mexico and Arizona. The Yucatan species is different from them, and as the case with many cave forms~ it differs in the longer appendages, legs, antennae, wings. Eremoleon longior n. sp. Body pale yellowish; head with a large black mark around antennae, truncate in front, above reaching to vertex; across vertex are three dark spots in a row, the laterals are transverse, the middle one consists of two elongate spots, close together and not divergent behind (in macer these diverge behind). Last joint of maxillary palpi partly dark, tip of antennae very dark; pronotum with a more or less distinct large dark mark, divided by a pale median stripe, each side behind are two shorter, curved pale stripes, uniting in front (much as in 1nacer); rest of notum largely dark, but with a median pale line on front part, and sorne pale each side near base of wings. Abdomen with segments dark on apical half above (as in macer). Legs pale, dark at tips of femora and tibiae and faintly toward base of tibiae. Fore wings hyaline, venation pale, marked with dark streaks (as in macer); many cross-veins wholly or partly dark; stigma white, with dark at base, and a larger dark cloud at rhegma, that near end of cubital fork scarcely distinct. Head as in E. macer; antennae much longer than in macer, reaching beyond base of the abdomen, latter a little longer th an in macer, pronotum also more slender than in macer. Legs and their parts also much longer th an in macer, the long inner spur of the front tibia being nearly as long as the distance from eye to eye. Wings a little longer than in macer, but with no more veins, the cells thus often longer than in that species; six to seven cross-veins before radial sector (eight or nine in macer), radial sector with eight branches, a few costals toward stigma are forked, the apical area is without gradate veinlets (normally several in macer). Length of fore wing 32 mm., width 9 mm. From Yucatan; Xtoloc Cenote Cave, Chichen Itza June 24; Chakxix Cave, Tekax, August 1; Puz Cave, Oxkutzcab, July 20; ail taken by A. S. Pearse. Type M. C. Z. No. 22654. Other specimens, which were doubtless E.longior, were seen in San Bulha Cave, Merida, J uly 13; Ebizt Cave and second cave on the San Roque Road, Oxkutzcab, July 18 and 23. Readily separated from E. macer Hag. by more elongate body, legs, and antennae; by few cross-veins before radial sector, and lack of apical gradate veins. According to the descriptions, Segura vitreus Navas from Cuernavaca, and N ovulga mexicana Navas from Vera Cruz are the same as Eremoleon macer; both are said to have eight or nine cross-veins before radial sector, and the figures show a denser venation th an E. longior. 1 have specimens of E. macer from Cuernavaca. 235

Bibliography of the Neuropterida Bibliography of the Neuropterida Reference number (r#): 93 Reference Citation: Banks, N. 1938 [1938.06.15]. A new myrmeleonid from Yucatan. P. 235 in Pearse, A. S. (ed.). Fauna of the caves of Yucatan. Publications of the Carnegie Institution of Washington 491:iii+1-304. Copyrights: Any/all applicable copyrights reside with, and are reserved by, the publisher(s), the author(s) and/or other entities as allowed by law. No copyrights belong to the Bibliography of the Neuropterida. Notes: File: File produced for the Bibliography of the Neuropterida (BotN) component of the Lacewing Digital Library (LDL) Project, 2015.