Ditomyiidae, Keroplatidae and Mycetophilidae) of Greece,

Transcription

1 CHANDLER et al.: Studia dipterologica 12 (2005) Heft 2 ISSN The Fungus Gnats (Diptera: Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae and Mycetophilidae) of Greece, its islands and Cyprus [Die Pilzmiicken (Diptera: Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae und Mycetophilidae) Griechenlands und seiner Inseln sowie Zypern4 1 by Peter J. CHANDLER, Dimitar N. BECHEV and Norbert CASPERS Mclksham (UK) Plovdiv (Bulgaria) Bechen (Gernlany) Abstract Key words ~ The spccics of fungu\ gnats (Bolitophilidae, Diadoc~dildae, Ditomyiidac. Keroplat~d:~e and Mycetophilidae) o~urring in Greece and Cyprus are reviewed. Altogether 201 species :Ire recorded, 189 for Greece and 69 for Cyprus. Of these 126 specie5 arc newly recorded fol. Greece and 36 arc newly recorded for Cyprus. The following new taxa arc described from Greece: Macrorrhyrtcha ibis spec. nov., M. pelargos spec. nov., M. laconica spec. nov., Macrocera critica spec. nov., Docosia cephaloniae spec. nov., D. enos spec. nov., D. pasiphae spec. nov., Megophthalmidia illyrica spec. nov.. M. ionica spec. nov., M. pytho spec. nov., Mycomya thrakis spec. nov., Allocolocera scheria spec. nov., Sciophila pandora spec. nov., Ryrnosia labyrinthos spec. nov.; M. ill\,ric,cr is also recorded troln Croc~lia. The following ncw taa are described from Cyprus: Macrocera cypriaca spec. nov., Megophthalmidia alrzicola spec. nov., M. cedricola spec. nov. The following neu synonymies are propod: M!,c,c~r~iw renuis I WXLKER, 1856) = M. interniissa PL.ASSMA~N, l984 syn. nov., Plrror~rtr ~1.illi.storri DLIFI>ZICKI, 1889 = P rnciscr CASFERS, 1991 syn. nov. A key is provided for thc western Palaearctic specie5 of M(ic-i.orrh~~~ic-Irci WI~~ERTZ. The composition of the fauna of thew Pamilies in Greece and Cyprus is discusscd in relation to the Mediterranean and to the wider dislrihution of genera and spccics. Diplcra, Sciaroidea, Bolitophilidae, Diadocid~idae. Ditornyiidac. Kcroplatidae, Mycetophilidac. faunistics, taxonomy, new species, key to hloc~rc~rrhync~/~ti bpecles, Greece, Cyprus Zusammenfhssung Vorliegende Publikation hat die in Griechenland und Zypcrn vorkomrnenden Pil~rniickcn (Bolitophilidae, D~adocidiidae. Ditomyiidae, Keroplatidae und Myce~ophil~dae) rum Gegen\tand. Insgesamt ucrdcn 201 Arten ge~neldel, 189 aus Gr~echenland und 69 auh Zypcrn. Von tliesen werden 126 erhtmals fiir d ~e Fauna Griechenlands und 36 erstrnals fiir die ~ypriolische Fauna rcglstrlert. Folgende neuc Artcn werden aus Griechenland bcschneben: Macrorrlzyr~cha ibis spec. nov., M. pelargos spec. nov., M. lacorzica spec. nov., Macrocera crilica Fpec. nov., Docosia cephalortiae spec. nov., D. erzos spec. nov.. D. pasiphae spec. nov.. Megophlhalmidin illyrica spec. nov., M. ionica spec. nov., M. pytho spec. nov., Mycornyu thrakis spec. nov., Alkocotocera scheria spec. nov., Sciophila pandora spec. nov., Ryrnosia labyrirtthos spec. nov.: hf. illyric.tr wird auch aus Krollt~en gerneldet. Folgcnde Arten wertien au\ Zypcr~i beschricbcn: Macrocera cypriaca spec. nov., Megophthalmidia alrlicola spec. nov.. M. cedricola spec. nov. Die nachfolgenden neuen Synonymc crgaben sich aua der Beclrbcitung des M;~teri;lls: hl\.c.onryu teriuis (WALKER, 1856) =M. ir~~ertnrssa PLASS\IANN. 19x4 syn. nov., Phl~)~~iti r~.illi.\rorti DLILDZI(.KI, 1889 = P. incisti CA~PERS, 1991 syn. nov. Es u urde ein Best~rnrnung.;. schliissel der westpal3arktischcn Artcn der Gattung Mtrc~rorrliyr~ch~r WIN~LRT~ erarhe~tel. Die Zu\arnrnenset~ung dcr Fnuna d~eser Familien in Gricchenland und Zypern wird iln Vcrgle~ch zu jener in dcr Mcditerraneih di\kutiert und crwcitcrt d ~e Intormationen lum Vorkommen dcr Gattungcn und Arten. Stichworter D~ptera, Sciaroidea, Bolitoph~lidae. Diadocidiidae, Ditomyiidae. Keroplatidae. Mycctoph~litlac, Faunist~k, Taxonornie. ncuc Arten, Beatin~mung\schlliscl von Mtrcrorrlryrrclrci. Griechenland. Zypcrn -..

2 CHANDLER et al.: The Fungus Gnats... of Greece, its islands and Cyprus Introduction Knowledge of the fungus gnat fauna of the Mediterranean subregion of the Palaearctic has been increasing in recent years with the appearance of national checklists for Italy (MIKEI.LI et al. 1995) and the Iberian peninsula (CARLES-TOLRA 2002), which have included the associated islands. Accounts of the species found in Israel (CHANDLER 1994) and Malta (CHANDLER & GATT 2000) have also appeared. At the same time checklists for this group have been provided for the Balkans (BECHEV 1997c including 338 species. of which 275 were listed for Bulgaria) and updated lists for Bulgaria (BECHEV 2002a and 2003: bringing the total for Bulgaria to 308 species and the Balkans to 356). BECHEV (l 997c) included a list of 27 species for the mainland of Greece. based on previous publications. Many of the additions reported here were included in the Fauna Europaea database con~piled by the first author (CHANDLER 2004) with an indication that they were new records to be published formally later. Information on the Greek fauna has been sparse in the literature and entirely based on collections made by visiting entomologists from other parts of Europe. The lack of any local collectors means that the coverage is of necessity patchy although collections are now available from several parts of the mainland and from most of the larger islands, so a distribution pattern. at least for the Mediterranean elements of the fauna is beginning to emerge. It is recognised that many further species will be added, especially with more collecting in the northern parts of the Greek mainland, where a similar fauna to Bulgaria can be expected. but it is desirable to bring together here what is known as a basis for further studies. It is also considered appropriate to include records for Cyprus as there have been several recent collections made there and much of the fauna is in common with that of the Greek islands. Earlier data on fi~ngi~s gnats of Greece is scattered in several publications (LOEW l869 and 1871 ; LUNDSTR~~M and 1917; MATIL-E 1969a and 1977b: PLASSMANN 1984; VAISAUEN 1984a; CASPERS 1991; CHANDLER 1991; CHANDLER Bi RIBEIRO 1995; BECHEV 1995, 1997a and 1997b: MARTINOVSKY 2001 ; CHANDLER & BLASCO-ZUMETA 2001 ; CHANDLER 2002; KLJRINA 2004). These papers together recorded 63 species from Greece. From Cyprus 34 species have been reported (GEORGHIOL 1977; VAISIZNEN l983a; CASPERS 1991; CHANDLER 1994; KURINA 2004), most of them without precise data. In the present work the total is brought to 189 species for Greece and 69 species for Cyprus, with an overall total of 201 species. For comparison with Cyprus, the total for Crete is 70 species. Material The material on which this study is based has come from several sources. Most of it has been collected since 1970 but some older material in museum collections has also contributed to the data. The material examined by the authors can be summarised as follows: (1) NC examined the large collcctions obtained from many parts of the Mediterranean region hy Dr H. M-\I.ICKI. in thc 1970s and 1980s. This included rnatcrial from many parts of Greecc and its islands, as well as from Cyprus. Turkcy and several parts of the westcrn Mediterranean. As described by CASPERS (1991) thc primary aim was to obtain Trichoptcra mostly by light-trapping but sornetimes by sweeping and collecting of non-target groups was incidental, with thc fi~ngus gnats in relatively IOW proportion comparcd to aquatic groups. Three spccies from Crecce and one from Cyprus werc dcalt with by CASPEKS (l 991). (2) DB cxarnincd collcctions made in mainland Grecce by I. SIVK and B. HORVAT (SMNH) in 1994 and by himself in the same year and in BECHEV (l 997a and 1997b) descl-ibed a ncw species and subspecies from this material. (3) PC cxamincd collbctions made by several British colleelors, principally Alan STI:RRS (1972 Corfu and adjacent mainland, 1979 mainland Grcece and the Pclopponesc Crete and 1982 Cyprus)

3 . Studia dipterologica 12 (2005) Heft 2: and Ian MCLEAN ( l980 Corfu Ccphalonia, 1982 Crctc and 1983 Cyprus), with s~nallcr amounts of material obtained on Corfu by John Is~iw, Milos by David H~NSHAW, Paxos by Tony IRWIN and thc Pclopponcsc by John KRAM~K and David GIBBS. Some records from this material have already bccn published by CASI'ERS (1991), BECHEV (1995, 1997b), CHANIII.ER & BI.ASCO-ZUMFTA (2001 ) and CHANIII-ER (2002). CHANDI-ER (1994) and CHANDLER & RIBEIR~ (1995) cited records of some specich as from countries or islands but without further detail. More rccently PC examined specimens collected on mainland Greece and Cyprus by Pa~ll GATT and Martin EBESER (Malta) and in scvcral parts of thc region from the collections of the Natural History Museum. London (BMNH), the MusCum National d'histoirc Naturcllc. Paris (MNHN), the MLIS~LII~ d'histoire-naturelle, Neuchztel, Switrcrland (MHNN) and thc Zoological Muscum. Copenhagen. Dcnmark (ZMUC) obtained by several other collectors. (3) Jan SEVC~K has determined material in the collection of the Silesian Museum. Opava (SMOC). This was mostly collected by Jindiich ROHACEK (Cyprus 2002 and Crete 2004), but also included some from the Greek ~nainland and Corfu, respectively collected by B. MOCEK and V. VRAREC. The localities from which material has been examined are listed below under the six regions recognised for recording purposes in the Fauna Europaea project. For mainland Greece and the Pelopponese the name of the province, followed by a colon, precedes the locality details and localities on the Greek mainland are listed under the six administrative regions. For these regions the Greek name is given followed by the English name in brackets. The collector's name is given following details of each site and if all the material with that data is located in an institution this is stated following the collector's name. For islands the usual English spelling is given first and where the Greek spelling differs this is given in brackets. The spelling of many Greek localities varies considerably between different maps, mainly due to differing methods of transliteration from the Greek alphabet. If the spelling on the latest maps consulted differs consistently from that on the label, this is given in square brackets. Depositories of Material examined BMNH MHNN MNHN NMBA SMNH SMOC ZMUC ZSM = The Natural Hi.;tory Museum, London. UK = Mu.;Cum d'histoire-naturelle. Neuchstel, Switlerland = MusCum National d'histoire Naturelle. Paris = Naturhistorischeh Museum der Bened~kt~ner Abte~. Admont, Auhtria = Slovene Muse~~m of Nat~~ral History, Ljubljana, Slovcnia = SlelhkC ~cmskc mulcuni (Silcsian Museum), Opava. Clcch Republic = Zoological Mu.;cum, Copenhagen. Denmark = Zoologische Staatsammlung. Munich, Germany GREECE - Marnland Greece and nearby rslnndh (Fauna Europaea area GR-GRC) MAINLAND: ~PEIROS (= Epirus) IPl. Preveza: Parga v.1972, olive groves (A. E. Srruss. BMNH) IP2. Preveza: Parga, Morfr Lake, 10-1 I.v.1972 (A. E. Sr~luss. BMNH) IP3. Preveza: Parga, Antho~~aha, 100 m v.1972 (A. E. Sr~uns, BMNH) IP4. Ioannina: Vikos Aoos National Park. Vikos, 1500 m, 23.v.2000, pasture, 23.v.2000 (B. MOCCK, SMOC) IPS. Ioannina: Pind Mth, Balndourna, Q~rc,rcus forest, 24.i (D. BLCHLV) IP6. Epir, Kokitos river, Themelo, (? probably 3Y014'0"N. 20"3l'5YvE) 40 In. 16.v.1994 (I. SIVFC & B. H01<1\1.) MAINLAND: MAKEDONIA (NW Greece = Greek province of Macedonia) MKI. Picrii: aouth of Plcri [= FtCsi]. 4.v.1979 (H. M\I I~KY) MK2. F16rina: Monastir, ~ir (5. GOLILULN, MNHN) MK3. Flcirina: Flcirina, \ii (H. M \R~FI.FT, MNHN) MK4. Flcirina: Flbrina, ii and iii.1918 (Dr Jovcrlx. MNHN) MKS. Flcirina: Zelova. near Flcirina. v. l9 18 (ARMCC DC L'ORICKT, MNHN) MK6. Flor~na: HolCvcn. ~oulh of Monaatir. ix-x.1917 (BI KICO, MNHN)

4 -p-p- CHANDLER et al.: The Fungus Gnats... of Greece, ~ts ~slands and Cyprus - p- MK7. F16rina: near HolCvcn, houth of Monastir, I918 (Dr BARBIFK, MNHN) MK8. F16rina: between Bukovo and Holiven, 1917 (BLINICO, MNHN) MK9. Thehsalonihi: Karasculi, near Sdonika I= Thcssalonikil. x.1917 (J. W.\TEKSI.ON, BMNH) MKIO. Thessaloniki: near Salonika I= Thessaloniki], (J. W~-~~KSTON. BMNH) MKI l. Fldrina: Pissoderi [= P~scrdiri m. l3.v. 1998, \vood and stream (P. GXIT) MKIZ. Flhrinn: Andartiko [= Ant:~rtikh], 1000 m, l3.~.1')~18 (P. GAI I) MK13. Pierii: Olyrnpus Mts, Litohoron [= Litochoro] vii.1953 (E. J ~ N S ~ & L N R. TOI.LLI.) MKI4. Kozinl: Picr~a Mts. easl of Velventos [= Velvendos], 40 14'05"N, 27"07'51"E, 1330 m, 19.\, (l. SI\'FC & B. HORV\T) MK15. Kozini: Pieria Mts, 1440 In, 19.v.1994 (1. SI\W & B. WORVAT) MK16. Kozini: Pieria Mts, 40"l 1 '.35"N, 22O05.3 I "E, , 19.v.1994 (I. SIVE~ & B. HOK~.AT) MK17. Drim:~: norlh of Sidironcro, 1190 m. 21.v.l994 (1. SI~LC & B. HOKVAT) MKIII. Drima: west of Sidironero. 4I023'l3"N, 24"12'03"E, 500 m. 21.v.I994 (I. SIVEC & B. HORV.\T) MK 19. Drdma: Ditihi Rotlopi. Elatia Forest, 1450 m. 21.v (1. SIVFC & B. HORV/\T) MK20. Dr;inla: Pfalacro Mts, norlh of Livaderh, 670 m. 21.v (1. SIVEC & B. HORVAT) klk2 I. Drima: north of Dipolama. IOClCl Irr, 22.v (1. Slv~c- & B. Ho~v,rj MK22. Drima: north of Dip61ama m, 4I023'32"N 24"38'46"E, 22.v.1994 (1. SIVEC & B. HOKVIT) MK23. Dri~rra: north of Diphtama, l030 m, 4In23'53"N 2J"3X'OhWE, 73.v.1994 (1. SI\EC & B. HOKV.AI) MK24. Drilnn: north of Dip6lanra m, 23.v (I. SIVFC & B. HORVAT) MK25. Drima: north of Diptitalna, 1400 m, 23.v.1994 (1. SIVFC & B. HORV~\T) MK26. Drima: south of Diptitama, 41"21'22"N, 24'.16'2OWE, 440 m, 23.v.lYY4 (I. SIVEC & B. HORV,\T) MK27. Pierii: Lilohoron [= Lithchorol. 40"06'02"N, 22"29'52"E. 250 m. Qrrrr.c.rrr ~.oc.c.ifi,-cr, 16.ik (D. BFCIILV) MK28. Thes\aloniki: Lagin~i, Pil~u., ~rijirtr-qrrc.~-c.~i.\ cocc,lfi.rir wood, I8.iv.1991 (D. BLCHLV) MK29. Kilkic: Go~~nrenissa, Qltr~-c.r~c-Ctrr~~irir~.\ forest, 26.i~ (D. BLCHEV) MAINLAND: THRAKI (NE Greece = Trakia, Thrace) TRI. ~vros: Lnutros, 14.1v.1981 (J.-P. H:\FY~I, MHNNI TR2. Rod6pi: Sapka Mts, Santla, 41 07'06"N, 25"49'linE. 220 m, 24.v.1994 (1. SIVLC & B. Ho~v.4~) TR3. Rodhpi: S;rpka Mts '07"N. 2S054'40"E, 545 m, 24.\,.1994 (I. Slvk(. & B. HORVAI) TR4. Rod6pi: Sapka Mts. 4I009'56"N, 2S055'I7"E, 735 m, 24.v (1. SIVF.I' & B. HOR~AT) TR5. Kod6pi: Anatoliki Rodopi, east of Dr~nli, 4I013'26"N. 25".?5'35"E, 240 m, 75.v.IL)94 (I. SILL(. & B. HOK~.AT) MAINLAND: THESSALIA (= Thessaly) TH I. Trikala: Pcrtuli I= Pcrtouli], 26.x. I972 (H. MAI.I(.KY) TH2. Trikala: 12 km cost of Pertuli [= Pertouli] (H. MAI.ICKY) TH3. bl:igni\ia: above Portalla ['! = Portariaj, 15.vi.1979 (H. MAI.ICKY) TH4. Magnisia: east of Chinia, 15.vi.1979 and 13.x.1980 (H. MALICKI ) TH5. Llirisa: Pierla hils, south of Livadi, 800 m, 18.v.1')94 (1. SIV~C. & B. HOKVA~) TH6. Lirisa: F;rr\al:r, IITM: FJ m. Pirrrrs 11igrcr forest, ZO.iv.1994 (D. Btc.~~cv) TH7. Trikala: Kastraki (near Kalamhaka and hle[cora). Querc,u.\-Cirr-l~irzlrs forest. 24.i~ (D. BECHF.\.) TH8. Magnisia: Pillon Malaki, 18.iv.1979 (M. DFTHI~R. MHNN) TH9. Lir~sa: Karia [= KnryA], 4O000'N/22"26E', 30.i~-l.v (H. M.ALI( KY) MAINLAND: STEREA ELLADA (= Central Greece) SEI. A~tolia-Akirnania (= Eloloakirnania): Agios Dimitrios, 25.x.1972 (H. MALIC~Y) SE2. Fokida: Mount Parnassos ahovc Polytiroson /= Polidrosos], 3.v.1976 (H. MALICKY) SE3. Fokida: 3 krn norlh of Kamhos. 450 m, v north facing deciduoua and evergreen oak \roods with stream (A. E. SIL.BBS) SE4. Fokida: 12 km soulh of Lidorihion [= Lidoriki], 500 m. v.1')79, stunted Querclc., ~.oc.c,(ferir (A. E. SI.L BB.$) SE5. Fohid:~: I k~rr west of Ap. Efthyrnia, 450 m, v.1979, small fields \vith trees (A.E. STLIBBS) SE6. Fokida: I km south-west of Amphissa [= Amfissa], 350 In. v.1979, olivc grove will1 stream (.A. E. S.ILBBS) SE7. Fokida: Delphi [= Delfi]. 5.v.1979 (A. E. SILIBBS) SEX. Fthihtida. Thcr~nopile I= Thermop!lcs]. hot spring and marsh. 14.v.1998 [M. J. EBPJER) SE9. Fokida: Drossohori [= Drosochhri]. Parnassos hlts, 800 m. 15.\..1998, conifer forcst (P. GATT) SEIO. Fokiil;~: Drossohor~ [= Drosochtiri], Parnassoh Mts, 850 m, 15.v.1998, dwarf oak and rneadows (P. GATT) SE1 I. Fokida: Eptalophos [= Epllilolos]. Parnassos Mts m, 16.v.19U8. meadows and conifcrs (P. GATT and M. J. EBEJFRI SE17. Fokida: Gravia. l6.v lneado\vs and riverhank [P. G.ATT) SE 13. Fokida: Gravia, UTM: FH28, 730 In. l.ix.198? (D. BECHFV) SE 14. Fokida: south of Kastel~a, Elconas. UTM: FH78, Q~lercus forest, 21.iv.1994 (D. B ~(.HE~ I

5 Studia dipterologica 12 (2005) Heft 2: MAINLAND: ATTIKI (= Attica) ATI. Attiki: Parnea, north of Athens [= Athina], 23.v (H. MALI(.KY) PELOPPONESE (PELOPONNISOS) 1. Arkadia: Alepochorion I= Alepochoril. 15.ix.1972 (H. M \I ICKY) 2. Argolida: Kek~larion [= Kek~liri], 18.x (H. MALICKY) 3. Lakonia: near Mistras I= Mystris], I3 km weat of Sparta I= Sparti] (H. M,u I ~KY) 4. Arkadia: caal of Tripolis L= Tripoli], 26.iv.1979 (A. E. STIIRHS) 5. Arkadia: Lake Takka. 27.iv.1979 (A. E. STI~HHS) 6. Arkadia: south-west of Lake Takka. 27.iv shaded slrearn from spring (A. E. SI I HHS) 7. Arkadia: north-west of Tripolis [= Tripoli], 28.iv.1979 (A. E. STLIBRS) 8. Arkadia: l km west of Vitina, 900 m, 28.iv.1979, grassland (A. E. STI:BB~) 9. Arkadia: 2 km north-west of Langadla. 800 m, 28.iv.1979, steep deciduous wood (A. E. STIIRH';) I0 Ileia: Olympia. l70 m, 29.1v.1979, dry woods, river and stream (A. E. STI~HHS) I I Acha~a: Kalogri;~. l.v coastal dunes with slack (A. E. STIIRRS) 12. Arkadia: near Vitina, Milaoundas. 900 m, 7.vil and 300 m, dry cave, 5.vili.1969 (L. Ts~c.~s, MNHN) 13. Arkadia: Vitina, Dracotrypia cave. 20.v1i. l969 (CARAYO~, TSA(.AS & PLIIOT, MNHN) 14. Achaia: (Mt) Hellnos, 1700 m. I0.vi. 1990, Ahie.s cepl~trlo~~icrr Corcst (J.-P. H ~~NNI Oi C. DLI~OUR, MHNN) 15. Lakoni;~: Taigc~os Mts, 8 km wsw oet11pi [= Trypi], 1100 m, l I.v. 1900, stream (J.-P. H~EKUI & C. DL~~OIIK, MHNN) 16. Korinthla: 6 km norlh of Kalivia, l600 m, 8.vi. 1990, Ahrrs c.c~phalor~ic.tr forest (J.-P. H,aruul & C. Drl~or R, MHNN) 17. Achaia: Helmos, 6km wsw of Ag. Varvlira, 1300 m, 9.vi.1990, caacadc (J.-P. H~LUUI Oi C. DLIIOI K, MHNN) 18. Lakonia: Taigetos M&, 7km waw oc Palopanayia, 200 m, 12.vi.l990 (J.-P. H.stuul Oi C. DVIOLK, MHNN) 19. Korinthia: Archeon Phencos, by River Erymanthios, 17.iv.2003, pinc forcal (J. KKARILK) 20. Korinthia: Archcon Pheneoa, 20.iv.2003, atream by monastery (J. KR~MLK) 21. Korinthia: Archeon Pheneos. by River Erymanthios, 26.iv.2003, near Lake Doxa L= Doxis] (J. KK \~IFK) 22. Achaia: Mt Chelmos l= Helmos], above Kalavrita [= Kalli\'ryta], 1700 m, vi.1982 (B. SKLL~ Oi S. L~UGL- MAKK, ZMUC) 23. Lakonia: 5 km aouth of Monemvasia. varlous dates in periods 10.xi-23.xii.l i.1084 and 17.v-25.v (G. CHKISTEYSEU, ZMUC) 24. Lakonia: Monemvaaia, 24.x. I984 (G. CHKIS~LNSLN, ZMUC) 25. Achala: near Koutia (E. JLIPIILL & V. ODIZIKE, MNHN) 26. Acha~a: pen day^, 38"35'N/22"00'E, 950 m, 2-3.vi.1975 (H. MALI(.KY). 27. Arkadia: Kastria, Ton Lirnnon cave. IS.v.1974 ("MIIs. GEYFVF", MNHN) 28. Arkadia: west of Vitina, 21.v (H. M~I.ICKY) 29. Korinthla: Archeon Pheneoa, 6.v.2004, stream by monastery (J. KRAMER) 30. Kor~nth~a: Archeon Pheneos. 7.v.2004, upper meadow stream (J. KK~MER) 31. Korinthia: Archeon Phcncoa, River Erymanthioa, 8.v.2004, ncar old Acfr trees (J. KR.~MLR) 32. Korinlhia: Archeon Phcncos, Lakc Doxa [= Doxia], 8.v.2004 (J. K KA~T~K) 33. Korinlhia: Archcon Phcncoa. Lakc Doxa [= Doxis], 9.v.2004, atream margin (J. KKAMLR) 34. Korinlhia: Archcon Phcncos. Lakc Doxa [= Doxia], 10.v.2004, slrcam margins (J. KRAMLK) 35. Lakonia: Mini, Stoupa sl., UTM S , 24.v.2004 (D. GIHHS) CEPHALONIA (KEFALONIA) 1. Lixouri. 14.v elm hedge by flowery meadow (I. F. G. M(.Lrku) 2. Mount Enos, 1100 m, 15.v.1981, l~rnestone hillside (I. F. G. M~LFAu) 3. Mount Enos, l000 m, 15.v.1081, dry gullcy (I. F. G. McLtk~) 4. Mount Enos \ummil, 1630 In, 15. l8 and 20.v.1981, Ahicjs cc~ph~ilorrico foreat (I. F. G. MCLLAN) 5. 1 km norlh-we51 of Kato Kalelios, l6.v.1981, dry gorge (I. F. G. McLt.sn) 6. Kato Katel~os, 16.v.1981, stream from road to sea shaded by plane tree5 (I. F. G. McLt.,k~) 7. Mount Enos, 1400 m, 18.v.1981, Ahivs cc,phulonicci forest (I. F. G. MCLFAN) 8. Pastra, 19 and 22.v wooded gorge with stream (I. F. G. MCLEAU) 9. North-weal of T~anata, I9.v.1981, ahaded stream with mossy seepages (I. F. G. M(.LE~u) 10. Mount Enos, 1300 m, 20 and 23.v.1981, area w~th large amount of dead wood (I. F- G. McLF~) 11. Mounl Enos, cast side, 2-4.vi.1977 (H. M~LICK)) 12. North-west of Taanala, 28.1~ (H. MALICKY) CORFU (K~RKIRA) 1. Ayia Kiriahi, 2.v.1072 (A.E. SII HHS, RMNH) 2. Dassia, 4.v.1972 (A.E. ST~HHS, RMNH)

6 CHANDLER et al.: The Fungus Gnats.,of - Greece, ~ts islands and Cyprus 3. Spartilas, 500 m, 3.v.1972 (A.E. Sr~nus, BMNH) 4. Strrnillas. 700 m. 3.v (A.E. Sr~'oss, Bh1NH) 5. mar bell;^ Beach Hotel. 5 km north of Mesmngli, 75.vi.1976 (J.W. 1 ~~4)) 6. Benitscs, 29.iv olive groves ulth rich ground flor;~, near stream1 (I. F. G. McLF.\N) 7. Benitccs. 29.iv.1980, beside stream in snlall gorgc (I. F. G. MCLLAN) 8. Kavos, l.v olive proves with rich ground flora (I. F. G. Mc'Lrm) 9. Kavos, I.v beside track to monastery (1. F. G. Mc'LrN) 10. Kavos, nlixed woodland around Inonastrry Panayia Arkoudillas (l. F. G. Mc LEA^;) I I. Gasto~lri, 4.v v.1980, narrow gorge sorlth of village (1. F. G. McLr:<N) 12. Palcocastritsa, 5.v.1980, dry gorge north of village (I. F. G. ~Z~.LF.AN) 13. Linia, 5.v.1080, track towards Lake Koris\ion, rich ground llora (l. F. G. McLE:\N) 14. Mount Pantokritor, 2 km wesl of main peak, l000 m, O.v.1980 (l. F. G. McLmr;) 15. Kcntromn. small caves bcside gorge to the south-west. 9.v (I. F. G. MCLEAU) 16. Kinopiaslcs. 11.v. 1980, roadside vegetation t I. F. G. M~LF-\N) 17..4g. Deka, l l.v.1980, dry gorge north of village (I. F. G. MCLLAN) 18. Ag. Deka to Bcnitses, 11.v olive groves (I. F. G. McLE.\N) 19. Ea\t of Tomploni. 10.~ (H. MALICKY) 20. Nesaria. l l.vi.l977 2nd 9.xi.1980 (H. MXLICKY) 2 1. Barbati, vi.2002 (V. VK~BFC-, SMOC) EUBOA (EVVOIA) I. Southwest of Plataniatos. lo.u.1980 (H. M~LICK)) 2. Ano Steni, 12.x.1980 (H. M-\LITKY) 3. Strni Dirfia, 4.vi (W. M-\LICKY) 4. Above Stropones, 24.v.1978 and 1 I-I2.x.1980 (H. ~~;\I.IC.KY) 5. South of Komitou, 6.vi.l979 (H. ~I-\I.Ic.K\) 6. Agios Dilnitrios v.1974 (H. M,ALIC.KY) PAXOS (PAXI) (near Corfu) I. Gaios. 12.v. 1980, damp valley among olives (A.G. IR\\IV) 2. Agrilas, 18.v. 1980, coastal scrub (A.G. IKWIN) SAMOTHRACE (SAMOTHRAKI) 1 Thernie [= rherm~], 16.\ (G1 IC-II-\RD & H ~R\ F?, BhINH) THASOS (THASOS) 1. Above Mal-iis, 17.vi.1979 and 17.u.1980 (H. M,\I.Ic~) km north-east ot MariCs, 17.v (H. MALICKY) 3. 5 km easl of MariCs. 17.x (H. M.~LIC.KY) 4. Dipotarnos, 1 8.vi (H. M ~LIC~Y) 5. Abovc Prinos. Ih.vi. l979 (H. M \I ICK)) North Aegean Islands (VOREION AIYAION) (Fauna Europaea area GR-AEG) CHIOS (KHIOS) 1. 2 km north of Fili, 28.v (H. M,\[ ICKY) 2. Pelinato rnount:~in range. altitude m, 17.v.1073 (H. MALICI(>) 3. 1 kni north of Spnrtunda, 22.v (H. MALIC~Y) 4. 5 km northeast ot Pirama. 20.v.1975 (H. MAI.ICKY) 5. South of Kurunla, 28.v.1973 (H. MALICKY) 6. North of Ker:~mos (H. M.\I ICKY) IKARIA (IKAR~A) I. Raches. 31.v. l979 (H. M.AIUCKY) LESROS (~,ESVOS) I. South ol' Neochorion. 25.v. l975 (H. M.ALICKY) 2. Arnbcliko, 75.v.1975 (H. Prl.41.1(.h\

7 Stud~a d~pterolog~ca 12 (2005) Heft hrn northwest of Agiabos, 20.v (H. M.\I ICKY) 1. l km norlhueat of Megolochiri, 24.v.1975 (H. MALICKY) 5. Myt~lcnc (= Mitilini), iv.1917 (Dr L\N~RIF.II. Arm& de I'Oricnt. MNHN) SAMOS (SAMOS) 1. Brlou Manolares, v (H. M ~LI(.KYI Cyclades Islands (KIKLADES) (Fauna Europaea area GR-CYC) ANDROS (ANDROS) l. Revmata. 21.X (H. MALI( KY) 2. Norlh of Apikia, 13.v (H. MALICKY) KITHIRA I. West of Avlemonas, 8.v.1976 (H. M~LICKY) MILOS (MILOS) I. hlala~be trap in Citrus orchard, v.IOX5 (D..!. nf C. Htus~.+w) NAXOS ( ~Axos) 1 Above Koron~\ X 1980 (H M~LICK) PAROS (PAROS) I. Northwe\t of Pctaludc\. 18.v (H. M ~LICKY) Dodecanese Islands (DODEKANISOS) (Fauna Europaea area GR-DOD) RHODES (RODHOS) I. West of Profitis Ilias, 3.iii.1982 (H. M.ILIIKY) 2. West ot Eleousa, 3.iii.1982 (H. MA Ir KY) 3. Bctkveen Psinthos and Arch~polis. 2.1ii.1982 (H. MALICKY) 1. 2 km north of Laerma, S.iii.1982 (H. h141 I ~KY) 5. 2 km norlhccrst of Profilia, 6.v.1975 (H. ~I.\LI~KY) 6. 1 km west of Kellilhia, lo.v.1975 (H. MALICKY) 7. 5 km north of Laerma, 3-l.iii.1982 (H. MALI~~) 8. Epta Piges, 4-6.x (H. M\LICKY) Crete (KR~TI) (Fauna Europaea area GR-KRI) 1. Gcorgioupoli\, 18.x1i.l981 (H. M.41 I~KY) 2 Kefalion, 16.v (H. ~I.III( KY) 3. Kakopetros, I.ix. 1972, 3.v ~. 1977, 13.v and 20.ii.1982 (H. M,\[ I ~ K Y ) 1. Gorge of Topolia, 20.v.1977 (H. MALI( KY) 5. East of Ierapetra. 18.i~ (H. MALI~KY) 6. Pantanasa, ix.1972 (H. M \LI(.KY) 7. Easl of Agios Ioannis. 8.\.1979 (H. MALICKY) 8. Mesa Pot~lmi. 8- I 0.v (H. MALIC.KY) 9. Kot\ifiana. 20.v (H. MAI.ICKY) 10. Monl Veniou. 23.v and 18.v (H. MALIC~) l I I. Fassa valley, weal of Chliaro, v.1977 (H. MALICKY) 12. Aligi near Siws, 2.x and 19.v.1977 (H. MALICKY) 13. Kat~omatados, 2-3.x (H. MALICKY) 11. Killaria ncar mouth, 18.ii (H. M.~LICKY) 15. Kalonichtis, 14.i~ and ii (H. RI.\l ICKY) 16. Karninaki (H. MALIC~) l 17. Near Kefali, 20.ii. l982 (H. MAI~ICKY) 18. Knossos, 150 m, and 2X.iv.1980, shaded area by htream and flowery lanes (A. E. STIIBES) 19. Schisrna, Elounda, 17.i~. 1980, saltnlarsh (A. E. Srrless) 20. Kritsa, 500 m. 18.1~ rochy valley cast of village (A. E. STL ooa)

8 -P P- p- p- -- CHANDLER et al The Fungus Gnats of Greece, ~ts Islands and Cyprus kni south of Paliilni. 500 m, 19.iv.1980, field by stream (.4. E. SI.IIHHS! 22. Gortyna,.Aghii Deka v.1980, field by stream (A. E. STLIBRS) krrr north of Mcla~nbes, 450 m. 22.iv by slre;iln (A. E. ST~HHS) km south of Spili, 500 m, 22.iv. 1980, Qur,n.~r.\ c.ocrfir-(i by slre:uii (A. E. STL us) 25. West side of Huni;~. 24.iv coastal dunes wilh river (A. E. S? cues) 26. Topolia, by Aghiu Sophia cave. 300 nl. 26.i1.l980, scruhhy hillside (A. E. S.r~.ues) km north-west of Siroules. 400 m. 26.iv shaded riivinc (A. E. Si i BBS) km north-east of Plakalonu, 27.iv scrubby hillside (A. F.. STLIHHS) 29. Hliaro, 200 m, 27.iv.1980, slrcam and orange grove (A. E. Srrlsss) 30. North of Rodopou. l3.v. 1982, i\olated tress and rock clefts (I. F. G. R.lcL~.\h I 31. South of Rodopou at Aspra Ner;r crossroad\, 13.v.1982 (1. F. G. Mc.LF~N) 32. Aspra Nera crossroads lo rn:lin road. 13.v.1982, irlive grebes ( I. F. G. Mc.Lr:\N) 33. Rodopou, 13.v. 1982, stream gorgc (I. F. G. M~LFAN) 34. Samaria Gorge. 14.v.IW2, pinc and cypress l'orest (I. F. G. McLr.4~) 35. Samaria Gorse. north of Saiiiaria Vill;~ge and spring south of village, l4.v.1982 (1. F. G. McLt-.4~) 36. Kotsifi;~na. 16.v shaded stream and olive groves (I. F. G. McLtm) 37. Near junction for Skord;ila, 17.v by olive grove (I. F. G. McLtm) 38. North of Omalo\, 1500 m, 17.v. 1982, small gorgc (I. F. G. Mc.LF.~N) 39. Samaria Gorge. 17.\.1952, 1 km in, sweeping Pirlrrs hr-urict (I. F. G. McLL.~N) 40. Ornalos Plate;lu. 17.v pond n1;lr:in (I. F. G. blrle.\~) 41. Near Askyfou, before ascenl to lnibros Plateau, 18.v.1982 (I. F. G. MCLFAI) 42. East Lassithi Plateau, 19.v. 1982, stream bed (pools only) (I. F. G. McLc.4~) 43. West of Mesa Polami. 19.v.19S2. flowing stream in cullivated area (1. F. G. McLt.4~) 44. Between Vuthipetro and Choudelsi. 20.v.lL)X2, almond tree (I. F. G. McLt~x) 45. Bclween Choudet\i and Ch~~rnbas, 20.v.1982, almond Lree (I. F. G. McLL%u) 46. Vai Pal111 Forest, 2 1.v (1. F. G. MC-LL.~) 47. Mount Psiloritis, in, 24.v. 1982, ascent to snowfield (l. F. G. McLlr,\~) 48. Mount Psiloritis, 1600 m. 24.v edge of snow patches (1. F. G. McL~tv) 49. Mount Psiloritis, Ideon Antron cave, 24.v.1982 (I. F. G. McLE.~~) 50. Mount Agion Pneuma In. 29.v.193h (R. E. G.~THOKNL-H.\III>Y, BMNH) S 1. SW Crcte, Imhros gorge. 12.v.1004, ;\long brook (J. ROH.\CLK, SMOC) 52. NW Crelc. Vrysscs. 3 km NW. 12.v.2004, mixed forest (M. V.iL.4, SMOC) 53. NW Cretc, Stilos. I km N. l3.v.2004, riverside. under trees (J. ROIIACIX, SMOC) 54. SW Crete. Samaria zorfc, NW part, 14.v.?004, under trees at hrook (J. RoII.~~.FK. SMOC) 55. SW Crelc, linhros gorge. southern part. 15.v under trees and rocks (J. ROHICLK, ShlOC) 56. NW Crete, Episkopi. Petres river. Ib.v.2004, under trees on ri\cr hanks (J. ROH\?FK, SblOC) 57. W Crcte. Omalos. 3 km SW. 17.v.2004, under lrces in pasture (J. RoH.\CEK, SbIOC) 58. W Crete, Prase\, 4 k ~n SW, 17.v along small brook in forest (J. KOIIACEK, SMOC) 59. W Crete. Prases. 4 km SW, IX.v.2004, ;11ong creek (J. ROH,ii.llK, SMOC) h0. W Crcte. Prases. 2 km W. 21.v ~~nder C<ist<~r~rci Lrees (J. RoH.ii.kti, SMOC) 61. W Crelc, Prases. 4 km SW. 21.v.2004, near small creek (J. Rot1 \C.LK, SMOC) (71. W Crete. Nea Rou~nata. 2 km NE, 21.v along bl-ook in forest (J. ROH \&K, SblOC) 63. W Crcte. Kakopetros, l km S along brook (J. ROHACEK. SMOC) CYPRUS (Fauna Europaea area CY) l. South of Asi~nu, 14.v (H. M.~LICKY) 2. East OS Platrr iv.1974 (H. ~I.%LICKI ) 3. Pedhoulis (Trood(r Mount~~ins). 4.iv. l983 (H. ~I.\I.IcKY) 4. Creek near Vavla. 6.iv.IC)83 (H. M.~LICI(Y) 5. 7 km south of b1iliku1-i, l.v. 1'174 (H. M ~LI(.KY ) 0. 4 km cast oskakopetria. 14.v.1974 (H. M\I.ICKY) 7. Moni Trooditissa i~ (H. M.~L.ICK\ ) 8. Agios Nikolaos, I l.v (H. M;\LI(.I\Y) km WNW of Apia Anna. 26.iv.lC)82. fulleys with pinc (A. E. Srr.eus) 10. Tsidha. l.\ (A.E. SIL nus) I I. Cedar Valley, l200 In, 2.v cedar. pine and oak fol.esl (A. E. S-r~ ens) 12. Kambos. 785 m. 3.v. 1082, cultiv:~ted ~allcy bottom w~th stream (A. F.. STI.HH\) km east of Kykko. 050 ni. 3.v pine lhrest (A. E. ST~IUHS)

9 pp -- Studia d~pterolog~ca 12 (2005) Heft 2: l km west of Pedhoulis, 1130 m, 3.v. 1982, alder flushes and orchards (A. E. SI.CHHS) km north ol' Pedhoulds, 1050 m. stream in orchards (A. E. STLIBBS) 16. Prodhromus, l500 In, 3.v.1982, pine forest (A. E. STLIBBS) k~n south of Prodhromus, 1400 In, 4.v stream with ~Tders (.A. E. STIIRRS) 18. P.rrlo Plntses, In, 4-5.v. 1082, stream with alders and woods (A. E. SI.CIHHS) 19. Ayloh Therapon. I h.v.1983, roadside scruh and abandoned cultivation lerraces (I. F. G. MrL~-\hj 20. Platres. I200 m, 2 l.v. 19x3, stream In Golden Oak forcs~ (1. F. G. MYLEA\) 21. Platres, v. 1983, beside and above Caledoni~~n Falls. Golden Oak forest and plane trees by thc slrcarn (I. F. C;. hfc.l~~x) 22. hlount Olyrnpus surnrnil In, 22 and 23.v.IYX3, Pi11~i.s ~zi,yt.(i (1. F. G. hl~.l~r\h) 23. Cedar Valley, cahtern arm, l l00 In, 22 and 2J.v.1983, Goldcn Onh forest (1. F. G. MYLLA.~) 24. Prodhromus, 1200 m. 24.v.1983, hpring and strcarn gulley (I. F. C;. M~LLAN) 25. Cedar V~lllcy, western arm, 24.v.1983, dry stream in Goldcn Oak forest (1. F, G. hft-lp.-\\) 26. Cedar Valley, 1 l00 m, 34.v.1983, sccpages on roadside bank between east and west arms of the \,alley (I. F. G. McLk-\u I 27. Aslnou, iii.1953 (P. A. BL:KTOK, BMNH) 28. Kyperounda, 1500 m, iii.1953 (P. A. BIIYIO~. BMNH) 29. Kyrcnin, iii.1953 (P. A. B~lx-rou. BhINH) 30. Sl;lvros rn. iii.1953 (P. A. BL'XTO~. BMNH) 31. Platres, v (C. F. J~NKIN~, BMNH) 32. Dhavios. I0.i~ (K. M. GLII~H,\RD. BMNH I 33. Mesa Potamis, Troodo5 Mts, 12.vii.1937 (G. A. M~v~ohrorlsr.\~~s. BhINH) 34. Plantanin, Foreat Station, 1500 m, 20.iv.1947 (G. A. M \\RO\I~C~SI.\KIS, BMNH) 35. Troodos Mls, 1300 m, 7.ix i'.' collcctur, BMNH) 36. Akrotiri, 3J036'N/32"58'E, 22.iv.2002, beach and \:lltrnarah (P. G.\l-I and M..I. ERULK) 37. Pl;lrlo Platrca, Caledonian Falls, 1250 m. 34"34'N/35"52'E. 25.iv.2002, pine I'c~rest, stream (P. G\TT) 38..Akrounta Forest, 74.iv.2002 (P. Gm) 39. Agio\ Nikolaos. 2 km north-east, 780 m. 25.iv.2002, rnaquis (P. G\T-r) 40. Pano Lefkclra, 290 In. 26.iv.2002, darn, maquis, streinn (P. GA~T) 41. Lernesw Forest. 3 knl nest Arakapas, h00 m, 29.iv.2002, atrcarn (P. G ~I-I) 42. Lelnesos Forest. Kellaki, 650 m. 29.1v.2002 (P. GATI) 43. Ornodos, Kissouaa, "40'N/32"4X'E. 27.1v.2002, stream, rniscd I'ol.est (P. GATT) 44. Lirnassol, 29.i\.7002, beach (P. GAI-I.) 45. Diar~zos \,alley. K~dasr, 245 m. 3J0J3'N/32"25'E, 23.iv stream and mixed foreat (M. J. ERLJLR) h ~n south of Zygi, 34"44'N, 32"44'E, 24.iv beach, atrcarrl. 7ii111oris, PI~rci,vi~ritr.\ (M. J. EBFIIR) 47. Akrounla, 12 km north. 640 m, 3-I"4X'N/3.I0O5'E. 2J.iv plne forest and Cist~is (M. J. ERLJLKI 48. Akrounta. 130 m, 34"JX'N/33"30'E, 24.iv rncndow. stream (M.J. EBLJLR) 40. Troodoa Mts, Caledonian Falla, 34"54'N/37"52'E, 25.iv.2002, pine forest (M. J. EBFIIIR) 50. Troodos Mth. Caledonian Falls, 1220 m. 34"36'N/32"58'E. 25.iv.2002 (M. J. EHFIFR) 5 l. Malia, Ki\housa, 550 m, 34"48'N/32"47'E. 27.iv.21002, rnived forest (M. J. EREJLR) 52. Kalo Chorio. 6 km aouth, 580 In, 24"49'N/33"00'E. 24.i\,.2002, pine forecl, stream, ~naquis (RI. J. EHFIFR) km aoulh-west of Kakopctria. I l.iv.2002, brook (J. Ro~iCek, SMOC) 54. Troodoa Mountain\, Troodos env., I l.iv pine forest (J. KOH>\~.LK, SMOC) k~n enat of Kykko. I2.iv.2002, pine forcal (J. Ko~iCek, SMOCl 56. Pane Pl;~tre\. C;llcdonian Falls. 12.iv.2002, under Irec\ ;It brook (J. ROI+;~~.FK, SMOCl km north-caal of Agioh Nikol;los. 13.iv slcppc fureat (J. ROH.\~FK. ShIOC) Systematic treatment Data are given in full in the case of new species and of other studied type material. For other material only the site numbers are listed (dates also if different dates are represented by the same site data), followed by numbers of specimens examined, under each of the geographical divisions recognised above. In the case of the material collected by I. SIVEC and B. HORVT the numbers anti sex of the specimens were not recorded. For each species the type of known distribution is indicated under Remarks. with reference to the ten categories listed in the concluding section on Discussion. With the exception of Neterotrich~i. for which there is currently no family assignment, all taxa are arranged alphabetically. The systematic arrangement follows that adopted by CHANDI.ER (2004)

10 p- CHANDLER et al : The Fungus Gnats... of Greece, rts islands and Cyprus Heterotricha Group Heterotricha takkae CHANDI.KK, 2002 'Qpe material. Holotype:6, GREECE: Pelopponese: south west of Lake Takka (site 6), 27.iv.1979, swept by shaded stream, Icg. A. E. STlitltls (BMNH). Paratypes:?. same data as holotype (BMNH);?. Crete (site 70). Kritsa. rocky valley, I 8.iv. 1980; i', Crete (site 231, 2 km north of Melambos, by stream, 22.iv. 1980: E. Crete (sitc 28). 2 km west of Plahalona. scrubby hillside. 27.iv.1980 (leg. A. E. ST~BBS, in PC collection). Remarks. Distribution type 6. CH\UI>L~K (2001) described this species from Greek niaterial and also recorded it from Switzerland and Italy. Material. GREECE: Mainland: MK16; MK25. Family BOLITOPHILIDAE Bolitophila (Bolitophila) cirierea MEICEN, 1818 Remarks. New to Greece. Distribution typc 2. This is a widespread Palaearctic species. with records from most parts of Europe. the C:rucasus and the Far East of Russia (Z.~ITZFV 1999b). Bolitophila (Bolitophila) saundersii (CURTIS, 1836) hlaterial. GREECE: Mainland: MK24: Pelopponcse, 20, 16: Crete: l, 2d d. Remarks. New to Greece. Distribution type 2. This is also widespread Palaearctic species (Z.\rrztv 1999b) and occur\ in North Africa (Algeria: BUKGFILLE-B.\[ AC~STO 1966) and Madeira (CIIANDLFR & RIBEIRO 1995). Material. GREECE: Mainland: MKl h Bolitophila (Cliopisa) pselrdohybrida LANDROCK, 1912 Remarks. New to Greece. Distribution Lype 2. Another widespread Palaearctic specie\. reaching the Far East of Russia (Z\IT~FV 1999h). Family DIADOCIDIIDAE Diadocidia (Diadocidia) ferruginosa (M~:IGEN, 1830) hlaterial. GREECE: Mainland: MK 16: MK22. Thasos: 1, 2d d. Remarks. New to Greece. Distribution type 2. This is the most widespread Palaearctic spccies of thc genus, frequent in Europe and recorded from Iran (L.~.<iov~.s & MATII F 1972). Material. GREECE: Mainland: MK17: MK25. Diadocidia (Diadocidia) spirzosula TOLI.ET, 1948 Remarks. New to Grccce. Distribution type 2. Also a widespread Palaearclic species, including the Caucas~rs (Joosr & PLASSLI \Uh 1976) and Allai (ZAIT~FV 1994). Material. GREECE: blainland: MKIS. Diadocidia (Adidocidia) valida MIK, 1874 Remarks. New to Greece. Distribution type 3. This species is widespread in Europe and also known from Azerbaijan (ZAITZEV 1994). Material. GREECE: Corfu: 21, 16. Family DITOMYIIDAE Symmerrrs annulatus (MEIGKN. 1830) Remarks. New to Greece. Dislribution type 3. This is a widespread species in Europe and has also been recorded from Azerbai.ian (ZAITZEV 1994).

11 p- -- Studia dipterologica 12 (2005) Heft 2: P Family KEROPLATIDAE Subfamily Keroplatinae Tribe Keroplatini Cerotelion strintutn (GMEI.IN, 1790) = I~rrecir~oi~ (F-\BRICILIS, 1775). prcocc. Material. GREECE: Mainland: SE1 2 I 8. Corfu: 12. I d. 1 9 : I 5, I X. Remarks. New to Grcccc. Distribution typc 3. This species is widespread in Europe, including Bulg:rr~a (BFCII~L 2002a) and is also recorded from Iran (R.l..\r~~r 1969b) and Azerbaijan (ZAIILL\ 1994). Keroplatus reautnurii (DIJFOUR, 1839) Material. GREECE: Mainland: IPI. I 9; Saniothrace: 1, 24 1 Remarks. Distribution type 3. This species was recorded as new to Greece, from Corfu, by KLIRIN \ (2004). It is w~despread in central and southern Europe. including Bulgaria (BFCHEY 2002a) and has also been recorded Sroni Morocco (M,\TIL~ 1986) and A~crbaijan (ZAITZFV 1904). Tribe Orfeliini Atltlemotz halidnyi (LOEW, 1871) Material. GREECE: Pelopponese: I I, 1 d. Ccphalonla: 1, 1 P. Corfu: 4. 1 'i : d d, 1?; 17. I d. blilos: I, 15 d J. 7PP.Crete:1~l.1d~.1?;20,1?;22,1$:26.Id;29.ld;31,5dd,13;12.4dd:33,2?9;1?;37.ld;41, 56, l d. Remarks. D~stribution type 7. LOEW (1871) described this species from Rhodes as well as Italy (Sic~l!). M \TII.L. (1977b) g;~\,e a record lrom Patmos. It is widespread in the Mediterranean and also known from the Canary Islands and Madeira (CIIAUDLER & RIHEIRO 1905). The female from Cephalonia was taken at flowers of Apiaccac. Material. GREECE: Cephalonia: 8, l 0. Zsot~eurotnyia czernyi (STROBI., 1909) Remarks. New to Greecc. Distribution typc 6. STROBL (1009) described lhia spccies from a Span~sh male. The Greek female agrees with Lhc description and was tentatively determined as c:errl\,i h! Lolc ~IATII.~, but findin: oi' males would be desirable for confirmation. Material. GREECE: Ma~nland: MK6. Crete: 4, l i'. Zsoneuromyin semirufa (MEIGKN, 1818) Remarks. New to Grcccc. Distribution typc 2. This specics is widespread in the Palaearctic, cornnion in Europc and reaching the Far East of Russia and Japan (ZAITZLV I990bi It has many synonyms, being variable in coloralion. Macrorrhyncha WINNEHTZ, 1846 This genus was characterised by MATILE ( who recognised sewn European spccies and provided a key for six of them (except for M. hreviro.siris (Lu~~srnohr. 1911) of which he had no1 exurnincd a male) and noted that the Japanese specics M. ezoerrsis (OK,ADA, 1938) alw helonged hcre. Since then eight l'urther Palaearctic species and a subspecies have been described (Wu & YANG BEC.HEV 1992b and l997h, ZAI'I'LEV CHANDLER 1994, CH,ANDLER & BI.ASCO-ZUMETA and KLIRINA 2004), while three further spccies are being dexribed from Japan by UESUGI (2005). Also PLASSVANN (1978) described Asindlrlum c.rem/~lum, which BECHEV ( 19978) showcd to belong to Mrzct-orr11j~nc.htr and CHAN- L)LER (1995) indicated that this was synonymous with M. c.ollurti (TOLLET, 1955), previously regarded as a synonym oc M. fluvu WINNERTZ. It is considered likely from ZAITZEV'S figure5 that M. gorodxoi,i ZAITZE~ is also synonymous with M. collurti. His other species, M. I~iipollz Z-\I.~ZEV, 1994, from the Far East of Russia (Sikhote-Alin) is distinct from other Palaearctic spccies in having the process bearing an apical seta, internal to the gonostylus, long and slcnder. The addition of another three species in the present paper bring\ the European total to fifteen. with one other species described from Israel. In Europc this has proved to he a predominantly Mediterranean genus, with most species apparently rather local in occurrence. At least seven \pecies are now known to occur in Creecc and the Bulgarian hunu comprises thrcc additional spccics (M. c~ollnrti, M. fluvcl and M. ~,elekrr BECHEV. 1997). In view of this increasing knowledge of Mac,rorr/i\.tic.l~a a key is provided to the known western Palaearctic spccics.

12 CHANDLER et al.: The Fungus Gnats... of Greece, its islands and Cyprus P -~ ~pp~ Key to western Palaearctic species of Macrorrl~yncha WINNERTZ l Mediotergite bearing setae Mediotergite devoid of setae Body mainly or entirely black with some grey dusting. Haltere black. Proboscis as long as head height. Male genitalia, Figs 5, 11-12; ovipositor, Fig M. ibis spec. nov. - Body at least partly yellow. Haltere yellowish brown Hind coxa widely brown at base. Mediotergite dark brown on disc. Mesonotum with three well marked dark stripes, more or less fused posteriorly. Abdomen with tergites all dark brown. [Male genitalia: MATILE 1975, Figs 13, M. aiicae M.ATII,E - Coxae and mediotergite yellow. Mesonotum yellow; stripes if present at most brownish yellow. Abdomen with tergites at least partly yellow Abdomen with some brown markings on disc of tergites 2-5, 6-7 all dark brown. Stem of median fork 4 times as long as radiomedial fusion. [Genitalia of both sexes: MATIL.E 1975, Figs , M..flava WINNERTZ - Abdomen entirely yellow. Stem of median fork little more than twice as long as radio- medial fusion. Only female known. [Ovipositor: MATILE 1975, Fig M. italica COST^) 5 Thorax brownish yellow without distinct darker markings: if stripes are present only slightly darker than ground coloration Thorax at least partly dark brown or black...: Proboscis as long as height of head. Abdomen mainly dark brown, with yellow basal markings on tergites 1-5. Male genitalia (Figs 1, 13-14) distinctive, with gonostylus bearing narrow elongate processes... M. laconica spec. nov. - Proboscis shorter than head height, at most a little longer than height of eye. Abdomen all dark or with some tergites yellow on apical margins Proboscis about as long as eye height. Male genitalia bearing a dense brush of black setae on dorsal lobe of gonocoxites. [Genitalia of both sexes: MATII.E 1975, Figs , 25; CHANDLER 1992, Figs M. rostrata (ZETTEHSTEDT) - Proboscis distinctly shorter, not more than half eye height. Male genitalia with dorsal lobe of gonocoxites with setae not in a dense brush or absent Abdomen with tergites 3-6 yellow npically. Costa extends halfway from R, to M,. Male genitalia (Figs 3, 7-8) with dorsal lobe of gonocoxites weakly developed but bearing.. strong setae; ovipositor, Fig M. brevirostris (LUNDSTR~~I) - Abdomen entirely dark brown. Costa extends a third of distance from R, to M,. Male genitalia with dorsal lobe of gonocoxites elongate but bare (Fig. 4); ovipositor, Fig. 10. I Male genitalia: CHANDLER & BL,ASCO-ZU~,IE.TA 2001, Figs M. gallica CHANDLER & MATILE 9 Mesonotum and abdomen uniformly black Mesonotum dark on disc but side margins more or less distinctly yellow or brownish. Haltere yellow Gonocoxites lacking dorsal process. Bridge of gonocoxites and tergite 9 medially constricted Gonocoxites with well developed blunt dorsal process. Bridge of gonocoxites and tergite 9 medially narrow but not markedly constricted l1 Haltere yellow. Gonostylus broader in lateral view with ventral lobe more developed. [Genitalia of both sexes: CHANDLER Figs M. grrichardi CHANI)I,ER

13 - Haltere with black knob. Gonostylus narrower r--- p-pppp Studia d~pterologlca 12 (2005) Heft 2: pp ~ in lateral view without ventral lobe developed. [Male genitalia: B~ctrrv 1992b. Figs ;... I. veleka BECHEV i 12 Haltere with black knob. Proboscis distinctly longer than head, practically twice as long as eye height. Dorsal process of gonocoxites more 1,-.. 'c. -- l slender and not extending beyond apex of gonostylus. [Genitalia of both sexes: MATILE 1975, Figs l l, ; male genitalia: BECHEV 1992b, 1 Figs M. geranias (LOEW) - i Haltere yellow. Proboscis about as long as head. Dorsal process of gonocoxites long and broad, extending distinctly beyond apex of gonostyl lus. [Male genitalia: KURINA Figs atticae K~JRINA 13 Proboscis longer than head, twice as long as eye height. [Male genitalia; BECHEV 1997b. Fig M. geranias thracica BECHEV, Proboscis not longer than head, usually little more than eye height A smaller species. wing length mm. Pleura and mediotergite brown. [Male genital L- lia: MATII-E 1975, Fig4 14, M. hispanica (STHORI,) Fig. Male of M~lc.,r,,.,.lry,rc./7~, ),,. - Larger species, wing length about 4.0 mm or ico spec. nov., lalcr-al vicw of ~onocoxites and more ~ ~ [ ~ ( ) ~ t ~ l ~ ~ ~. 15 Vein R, meets costa less than twice its length from tip of R,. Proboscis longer than eye (associated female has thorax all dark so would run to couplet 10 above) Ple~lra and mediotergite all dark. [Genitalia of both sexes: CHANDLER 1994, Figs 26-29] ardea CH~NDI,ER - Vein R, meets costa more than twice its length from tip of R,. Proboscis at most as long as eye height. Pleura or mediotergite often with yellow markings Side margins of mesonotum distinctly yellow and pleura with yellow markings on mesepimeron and laterotergite. Proboscis as long as eye height. Male genitalia (Figs 6, 15-16) with gonostylus apically blunt with short dorsal processes; ovipositor Fig M. pelargos spec. nov. - Side margins of'mesonoturn obscurely brownish; mediotergite narrowly yellow laterally. Proboscis about as long as two-thirds eye height. Male genitalia (Fig. 2) with gonostylus bearing an elongate slender posterior process IMale genitalia: BECHEV l992a. Figs M. collarti (TOLLII.) Macrorrhyncha atticae KIIKINA, 2004 Remarks. Distribution type 10. Kt rclu~ (2001) dcscribcd this specie\ irom a sinsle male c~,llccted in thc ~lia~nl:~ntl Gr-eck PI-ovince of Attiki.

14 CHANDLER et al.: The Fungus Gnats... of Greece, its islands and Cyprus Figs 2-6: Male genitalia of Macrorrhyncha species, lateral view of gonocoxites and gonostylus: - 2: M. collarti (TOLLET). - 3: M. brevirostris (LUNDSTROM). - 4: M. gallica CHANDLER & MATILE. - 5:' M. ibis spec. nov. - 6: M. pelargos spec. nov. Macrorrhyncha brevirostris (LUNDSTR~M, 1911) (Figs 3,7-9) Asindulum brevirostre LUNDSTROM, l91 l (brevioste also given as name in error) Macrorrhyncha brevirostre: MATILE, 1975 (spelling not agreeing in gender with genus) Material. GREECE: Cephalonia: l l, 16 ; 12, 16. Paxos: 2, 16. Male. Head with frons yellowish brown, face yellow. Antenna shorter than thorax, with median flagellomeres less than twice as long as broad; scape, pedicel and base of first flagellomere yellow, antenna otherwise brown. Proboscis short, a quarter height of head. Palpus brown. Thorax entirely brownish yellow, with three stripes on mesonotum brighter and more orange. Acrostichals biserial; dorsocentrals irregularly triserial, situated between stripes. Some posterior prospiracular setae present. Otherwise pleura and mediotergite bare. Wing yellowish, unmarked. Vein Sc faintly reaching costa. Vein R, ending a little more than its length from tip of R,. Costa extends 0.4 distance from R, to M,. Vein An fading apically, faintly reaching wing margin. Radiomedial fusion setose ventrally. Haltere yellow. Wing length mm. Legs yellow, with tarsi appearing darker. Fore metatarsus 0.8 length of tibia. All setae and tibial spurs dark. Abdomen with tergites 1-2 yellow, 2 faintly brown near base dorsally; 34 brown on basal half dorsally, the dark area contracted laterally; 5-6 progressively more broadly brown, 7 narrowly yellowish on apical margin and 8 entirely brown. Sternites 1-3 yellow, 4-5 faintly brown at base, 6-9 marked as corresponding tergites. Genitalia (Figs 3, 7-8) yellowish brown; gonocoxites joined by a broad ventral bridge and with a rounded setose lobe dorsally.

15 Studia di~teroloaica 12 (2005) Heft 2: Figs 7-8: Male genitalia of Macrorrhyncha brevirostris (LUNDSTROM): - 7: Ventral view of gonocoxites and gonostylus. - 8: Dorsal view of tergite 9 and cerci. Female. Ovipositor (Fig. 9) is drawn from a Croatian specimen (BMNH). The ovipositor of M. gallicn CHANDLER & MATILE is figured (Fig. 10) for comparison. Remarks. New to Greece. Distribution type 8. The type locality (Novi, stated to be in Hungary) was in Yugoslavia (Serbia) according to KRIVOSHEINA & MAMAEV (1988) and it has also been recorded from Croatia (COE 1962b; material in BMNH). MATILE (1975), however, considered a female from Hungary to be conspecific. The record from Corsica by MATILE (1977a) was based on M. gallica so it has only been confirmed from south-west Europe. LUNDSTR~M (1911) described this species from a single male and the material examined agrees well with the genitalia figures given by him. The specimens described here also agree with his description, repeated by MATILE, except in the entirely yellow halteres (brownish on the knob in the type) and in the extent of the abdominal markings (tergite 2 brown basally and 3-6 mainly brown with narrow yellow hind margins in the type). Macrorrhyncha geranias (LOEW, 1869) Macrorrhyncha geranias thracica BECHEV, 1997 Type material. (subspecies thracica) Holotype: c?, GREECE (site TR2), Rod6pi: Sapka Mts, Sanda, 4I007'06"N, 25"49'43"E, 220 m, 24.v.1994, leg. I. SIVEC & B. HORVAT (SMNH). Paratypes: 16, l?, same data as holotype. Other material. GREECE: Milos: 1,7c? c?, 19. Remarks. Distribution type 10. L ~EW (1869) described this species from Rhodes. The male lectotype and female paralectotype were described and figured by MATILE (1975). BECHEV (1997b) described a subspecies thracica from mainland Greece (Thriki), which differed in coloration of the thorax, having the lateral margin and humeral area of the mesonotum and haltere yellow while the type material had these all dark. The Milos material agrees with the type in this respect. Macrorrhyncha guichardi CHANDLER, 1994 Type material. Holotype: c?, CYPRUS (site 32): Dhavios, 10.iv.1971, leg. KM. GUICHARD (BMNH). Paratypes: 16, same data as holotype. Remarks. Distribution type 9. The type material of M. guichardi was from Cyprus. CHANDLER (1994) also recorded it from Israel and figured the genitalia of both sexes, the female from Israeli material. The male genital structure is closest to M. veleka Bechev, described from Bulgaria.