Mosquito surveillance programs have been implemented

Size: px
Start display at page:

Download "Mosquito surveillance programs have been implemented"

Transcription

1 VECTOR-BORNE AND ZOONOTIC DISEASES Volume 17, Number 3, 2017 ª Mary Ann Liebert, Inc. DOI: /vbz Records of Aedes albopictus (Skuse, 1894) (Diptera; Culicidae) and Culex tritaeniorhynchus (Diptera; Culicidae) Expansion in Areas in Mainland Greece and Islands Eleni Patsoula, 1 Stavroula Beleri, 1 Annita Vakali, 2 Danai Pervanidou, 2 Nikolaos Tegos, 1 Andreas Nearchou, 3 Demetrios Daskalakis, 4 Spiros Mourelatos, 5 and Christos Hadjichristodoulou 6 Abstract Annual entomological surveillance programs aiming to monitor mosquito populations and record presence and absence of mosquito species have been performed in Greece. We report, in this study, new records and expansion of Aedes albopictus in the islands of Lesvos (region of North Aegean), Crete (region of Crete), and the regional units of Rodopi in East Macedonia-Thrace. Furthermore, Culex tritaeniorhynchus was recorded for the first time in Arta (region of Epirus) in northwestern Greece. Keywords: Aedes albopictus, Culex tritaeniorhynchus, entomological surveillance, pathogen vector, Greece Introduction Mosquito surveillance programs have been implemented on an annual basis in Greece, since 2010 when the West Nile virus (WNV) epidemic was recorded for the first time in Northern Greece. From 2010 to 2013, these programs were performed by the Public Health authorities, the regions, and the MALWEST program (Marka et al. 2013, Patsoula et al. 2016). During 2014 and 2015 transmission periods, entomological surveillance activities were carried out only in small scale, with the participation and cooperation of the subcontractors for the vector control programs. These programs were funded by regions/regional units throughout Greece and local public health authorities, in collaboration with the National School of Public Health (NSPH) and the Hellenic Center for Disease Control & Prevention (HCDCP). One of the main objectives of entomological surveillance is to record presence/absence of mosquito species, monitor mosquito populations and detect the geographical distribution, and possible expansion of mosquito species in new areas. Aedes (Stegomyia) albopictus (Skuse, 1894), the Asian tiger mosquito, is considered to be the most invasive mosquito species in the world (Gratz, 2004). Ae. albopictus first appeared in western Greece, in 2003 (Samanidou-Voyadjoglou et al. 2005, Patsoula et al. 2006) and is now established in the capital city of Athens (Attica region), and in many regional units in Central Macedonia, Thessaly, and Peloponese (ECDC VBORNET 2016, Giatropoulos et al. 2012a,b, Giatropoulos 2013, LifeConops 2015). Worldwide, it is important for public health since it is known to transmit arboviruses, such as Dengue, Chikungunya, and Yellow fever. It is also a vector of filarial nematodes, Dirofilaria spp. (Cancrini et al. 2003a,b, Gratz 2004, Becker et al. 2010, and ECDC 2012). In recent years, cases of Dengue and Chikungunya fever have been recorded in Europe (Marchand et al. 2013, INVS 2014, Paty et al. 2014). Culex(Culex) tritaeniorhynchus Giles is a part of the Culex vishnui subgroup (Toma et al. 2000), distributed throughout the Asian region, extending into the Middle-east, the Mediterranean and Afrotropical region, China, Russia, Japan, Korea, Indonesia (Lee et al. 1989), Northeast Africa, India, Iran, Iraq, Israel, Jordan, Lebanon, Maldives Islands, Mozambique, Saudi Arabia, Sri Lanka, Syria, Turkey, and Turkmenistan (Walter Reed Biosystematics Unit). In Europe, it has been reported in Albania (Danielovi and Adhami 1960, Adhami 1987) and Greece since 2003 from samples originating from a coastal marsh in the area of Marathon in the Prefecture of Attica (Samanidou and Harbach 2003). It has also been recorded in rice fields in the regional unit of 1 Department of Parasitology, Entomology and Tropical Diseases, National School of Public Health, Athens, Greece. 2 Hellenic Center for Disease Control and Prevention, Athens, Greece. 3 Andreas Nearchou Co-operative NOVA FARM SA Geotechno Ygeionomiki O.E, Xanthi, Greece. 4 Igionomiki Kritis-Daskalakis Demetrios, Chania, Crete. 5 Ecodevelopment SA, Filyro Thessalonikis, Thessaloniki, Greece. 6 Department of Hygiene and Epidemiology, University of Thessaly Medical School, Larissa, Greece. 217

2 218 PATSOULA ET AL. Etoloakarnania, Western Greece, from 2008 to 2013 (Lytra and Emmanouel 2014). Cx. tritaeniorhynchus is a vector of Japanese Encephalitis ( JE) (Bram 1967, Self et al. 1973) and isolates of other viruses, including Sindbis and Dengue, have been found in this species (Lee et al. 1989). Our findings reflect the expansion and further establishment of these two species in new areas of the country, derived from the national entomological surveillance programs. Materials and Methods Entomological surveillance was carried out from May to November (period varies per regional unit) and included placing traps for adult mosquito collections and identification of mosquitoes up to species level (Patsoula et al. 2016). The results from the entomological surveillance activities were directly communicated to all the involved local authorities (regions and regional units, Departments of Public Health and Social Welfare, Directorates of Sanitation Control). Traps for adult mosquito collections were placed in rural, urban, and periurban sites every 15 days, in 28 regional units in 2014 and 19 regional units in In 2014, a median number of 54 (range: 7 68) traps (CO 2 and Triple traps) were placed throughout the study period in the 28 regional units, while in 2015, a median number of 8 (range: 0 31) traps were placed in the 19 regional units. The entomological surveillance areas were chosen after evaluation of the existing data for vector-borne diseases in humans and animals from previous transmission periods. However, it was not possible to conduct the entomological surveillance in all areas recommended by the entomological protocol. The surveillance was carried out by the contractor companies, which were awarded tenders for environmental applications and mosquito control in the regions. The traps used were either CO 2 and/or Triple traps according to the contractors availability of the traps. The trap locations are presented in Figures 1 and 2, for the years 2014 and 2015, respectively. Traps were placed at dusk from *6:00 pm and were collected the following morning at 9:00 am. The contents of the whole trap were placed in a plastic container on dry ice and sent to the Laboratory of Medical Entomology in NSPH for identification. All mosquitoes were identified to the species level using morphological characters (Darsie and Samanidou- Voyadjoglou 1997, Harbach 1998, Samanidou and Harbach 2001, 2003, Schaffner et al. 2001). Ae. albopictus and Cx. tritaeniochynchus adults, which were collected for the first time in areas of the country, were also verified by polymerase chain reaction (PCR) methods. DNA was extracted from one leg of individual adult mosquitoes following the protocol described in Patsoula et al The nuclear ribosomal spacer gene ITS2 region of the rdna gene was amplified by PCR using 5.8S and 28S FIG. 1. Sites of adult mosquito collections, 2014.

3 RECORDS OF A. ALBOPICTUS AND Cx. TRITAENIORHYNCHUS EXPANSION IN GREECE 219 FIG. 2. Sites of adult mosquito collections, primers (Porter and Collins 1991, Wesson et al. 1992, Collins and Paskewitz 1996) and a region of the mitochondrial cytochrome oxidase I gene (COI) was also amplified using primers C1-J-1718 and C1-N-2191 (Simon et al. 1994). PCR was carried out with the protocol and cycling conditions as described in Patsoula et al The PCR products were run on 2% agarose gels. Gels were stained with ethidium bromide and bands were visualized under UV transillumination. Aedes albopictus products were subjected to a restriction fragment length polymorphism assay (RFLP) for further verification of the PCR results (Patsoula et al. 2006). PCR products for both genes were cleaned using a commercially available kit (DNA isolation spin kit agarose, Applichem, Germany), according to the manufacturer s instructions to be analyzed by sequencing. Results A total of 47,039 mosquitoes were collected in 2014, belonging to 14 mosquito species (Culex pipiens s.l., Culex theileri, Aedes caspius, Aedes dorsalis, Aedes albopictus, Aedes detritus, Aedes vexans, Culiseta annulata, Culiseta longiareolata, Anopheles sacharovi, Anopheles hyrcanus, Anopheles maculipennis s.l., Anopheles superpictus, Anopheles claviger). In 2015, 2183 mosquitoes were collected. They belonged to 13 mosquito species and 1 genus (Culex pipiens s.l., Culex tritaeniorhynchus, Aedes caspius, Aedes dorsalis, Aedes albopictus, Aedes detritus, Aedes vexans, Culiseta annulata, Culiseta longiareolata, Anopheles sacharovi, Anopheles maculipennis s.l., Anopheles claviger, Uranotaenia unguiculata, Coquillettidia spp.). In the context of the entomological surveillance for the years 2014 and 2015, Culex pipiens s.l. was the most abundant species (89% and 66.9%, respectively) and was collected in all study areas. During the entomological surveillance programs, in 2014 and 2015, Ae. albopictus populations were recorded in areas with no previous official records of the species. In addition, Ae. albopictus has been reported in areas where it has been sporadically found and has been established by now (ECDC VBORNER). Ae. albopictus has been detected for the first time in the regional unit of Chania in Crete for two consecutive years, in the island of Lesvos in the North Aegean region, and in the regional unit of Rodopi in East Macedonia & Thrace region. The species has already been introduced and further records were performed in the regional units of Halkidiki, Pieria, and Kilkis in the region of Central Macedonia, and in the regional units of Drama, Xanthi, in the East Macedonia & Thrace region. In addition, Cx. tritaeniorhynchus mosquitoes were recorded for the first time in the regional unit of Arta (Epirus region). In Tables 1 and 2, dates of mosquito collections, region, regional unit and municipality, the trap type that was used, environmental site, longitude and latitude, and the number of adult mosquitoes captured/trap/night are presented for Ae. albopictus and Cx. tritaeniorhynchus, respectively. The geographical distribution of mosquito traps that collected

4 Table 1. Data of Expansion of Ae. albopictus in Greece, 2014 and 2015 Date Region/regional unit Municipality Trap type Environmental type Longitude Latitude Number of adults captured/gender July 28, 2014 Central Macedonia/Halkidiki Kassandra CO 2 Natural, rural \Ae. albopictus August 4, 2014 Central Macedonia/Kilkis Kilkis CO 2 Periurban \Ae. albopictus, 2_ Ae. albopictus August 27, 2014 Central Macedonia/Halkidiki Poligyros CO 2 Natural, rural \Ae. albopictus November 18, 2014 Crete/Chania Apokoronou Triple Periurban \Ae. albopictus September 17, 2014 Central Macedonia/Kilkis Kilkis CO 2 Periurban _ Ae. albopictus October 13, 2014 Central Macedonia/Kilkis Peonia CO 2 Periurban, rural _ Ae. albopictus June 24, 2015 North Aegean/Lesvos Kalloni CO2 Marsh \Ae. albopictus August 18, 2015 Crete/Chania Chania Triple Periurban \Ae. albopictus, 2_ Ae. albopictus September 8, 2015 Crete/Chania Chania Triple Urban \Ae. albopictus November 3, 2015 Crete/Chania Chania Triple Periurban \Ae. albopictus, 1 _ Ae. albopictus July 23, 2015 East Macedonia &Thrace/Drama Drama CO 2 /dry ice Urban \Ae. albopictus August 11, 2015 East Macedonia &Thrace/Drama Drama CO 2 /dry ice Urban \Ae. albopictus August 11, 2015 East Macedonia &Thrace/Xanthi Xanthi CO 2 /dry ice Urban \Ae. albopictus East Macedonia &Thrace/Drama Drama CO 2 /dry ice Urban \Ae. albopictus September 11, 2015 East Macedonia &Thrace/Drama Doxato CO 2 /dry ice Urban \Ae. albopictus September 11, 2015 East Macedonia &Thrace/Drama Prosotsani CO2/dry ice Urban \Ae. albopictus September 11, 2015 East Macedonia &Thrace/Rodopi Komotini CO 2 /dry ice Urban \Ae. albopictus September 11, 2015 East Macedonia &Thrace/Xanthi Kvdira CO 2 /dry ice Urban \Ae.albopictus October 6, 2015 Crete/Chania Chania Triple Periurban _ Ae. albopictus 220

5 RECORDS OF A. ALBOPICTUS AND Cx. TRITAENIORHYNCHUS EXPANSION IN GREECE 221 Date Table 2. Data on Cx. tritaeniorhynchus Mosquitoes in the Prefectural Unit of Arta, Prefecture of Epirus, Western Greece, 2015 Region/ regional unit Municipality Trap type Environmental type Longitude Latitude Number of adults captured August 24, 2015 Epirus/Arta Nikolaos Skoufas Triple Urban \ Cx. tritaeniorhynchus September 4, 2015 Epirus/Arta Nikolaos Skoufas Triple Urban \ Cx. tritaeniorhynchus September 5, 2015 Epirus/Arta Nikolaos Skoufas Triple Urban \ Cx. tritaeniorhynchus mosquitoes of the aforementioned species in different areas is presented in Figure 3. PCR amplification using both primer pairs (5.8S/28S and C1-J-1718/C1-N-219) produced fragments of the expected sizes, specific for Ae. albopictus. The RFLP assay verified that all samples examined were Ae. albopictus. Sequencing results of the four ITS2 and COI products revealed sequences with % homology to the GenBank deposited ones for Ae. albopictus. One COI and four ITS2 sequences have been submitted to the EMBL sequence database, with the accession numbers LT and LT , respectively. Sequencing of the two Cx. tritaeniorhynchus PCR products, examined for both genes, produced sequences sharing 95 99% homology with GenBank deposited ones. The two COI fragments share 99% similarities with GenBank sequences with accession numbers KC and KC753197, regarding Cx. tritaeniorhynchus samples collected in rice fields in Etoloakarnania, Greece (Lytra and Emmanouel 2014). Two COI and two ITS2 sequences have been submitted to the EMBL sequence database, with accession numbers LT and LT , respectively. Discussion and Conclusion Since the WNV outbreak in Greece in 2010 and the cases of autochthonous malaria due to Plasmodium vivax reported in Evrotas, Lakonia, there is increased concern among Public Health authorities, regions regarding vector-borne diseases. Entomological surveillance programs are implemented to promptly monitor mosquito species presence/absence, detect pathogen species, and support mosquito control actions. Our findings confirm the suspicion that Ae. albopictus has been expanding in Greece, a fact that increases the possibility for further local transmission of viruses transmitted through Ae. albopictus after their importation. Ae. albopictus is known to be an effective vector of Dengue, Chikungunya, FIG. 3. Geographical distribution of collection sites of Ae. albopictus and Cx. tritaeniorhynchus in new areas of the country. The yellow pins represent sites where Ae. albopictus adults were collected and red pins the sites of Cx. tritaeniorhynchus collections.

6 222 PATSOULA ET AL. Yellow fever, and other viruses (Gratz 2004, Becker et al. 2010, ECDC 2012). The species was involved in local transmission of cases of Dengue and Chikungunya in Europe (Marchand et al. 2013, INVS 2014, Paty et al. 2014). These facts along with the severe nuisance caused by Ae. albopictus to residents of cities and villages should be of great concern to public health authorities. In addition, we report about the third record of Cx. tritaeniorhynchus in Greece in the Municipality of Nikolaos Skoufas, regional unit of Arta, region of Epirus, in Western Greece. Cx. tritaeniorhynchus was the most abundant species in this regional unit. These findings are important as they refer to urban settings and concern a regional unit in close proximity with Etoloakarnania where the previous records of the species were performed. It may suggest that the species has spread along the western part of the country. Cx. tritaeniorhynchus is also present in Albania (Danielovi and Adhami 1960, Adhami 1987), therefore we could hypothesize that the species is present and more widely distributed in western Greece. It has possibly been underestimated until now, since no data were available. Cx. tritaeniorhynchus is mainly distributed in south Asia, Middle East, and Afrotropical countries. It is a potential vector of important diseases and pathogens such as JE, Dengue, Rift valley fever, Sindbis, and microfilariae (Bram 1967, Self et al. 1973). This present record along with the previous ones indicate the need for further studies on Cx. tritaeniorhynchus distribution in other regions of the country. A limitation of the study was the geographical distribution and representativeness of the mosquito collection sites, as surveillance activities were mainly based on the voluntary participation of local subcontractors of the vector control programs. Thus, it was not possible to conduct entomological surveillance in all areas recommended by the protocol and only distribution data on presence/absence of mosquito species are provided. Currently, there is an increasing concern in Europe regarding vector-borne diseases (Semenza and Zeller 2014). The introduction and establishment of these two species impose risks for Greece, related to the pathogens that they can transmit in humans. This fact should be taken into consideration by public health authorities to implement integrated mosquito management programs, perform risk assessment, and work toward the development of a national strategic action plan, aiming to the prompt management of diseases transmitted by invasive mosquito species. Acknowledgments The authors would like to thank all regions and regional units and the subcontractor companies for mosquito management that performed adult collections and sent the samples to the Laboratory of Medical Entomology of NSPH, for participating in the Entomological Surveillance programs of 2014 and Author Disclosure Statement No competing financial interests exist. References Adhami J. Mushkonjat (Diptera: Culicidae) te Shqiperise, tribu Culicini. Revista Mjekesore 1987; 1 2:82 95 (in Albanian). Becker N, Petrić D, Zgomba M, Boase C, et al. Mosquitoes and Their Control. DOI: / , 2nd edition. Berlin Heidelberg: Springer-Verlag, Bram R. Contributions to the mosquito faunaof Southeast Asia. II. The genus Culexin Thailand(Diptera: Culicidae). Contrib Am Entomol Inst 1967; 2: Cancrini G, Frangipane di Regalbono A, Ricci I, Tessarin C, et al. Aedes albopictus is a natural vector of Dirofilaria immitis in Italy. Vet Parasitol 2003a; 118: Cancrini G, Romi R, Gabrielli S, Toma L, et al. First finding of Dirofilaria repens in a natural population of Aedes albopictus. Med Vet Entomol 2003b; 17: Collins FH, Paskewitz SM. A review of the use of ribosomal DNA (rdna) to differentiate among cryptic Anopheles species. Insect Mol Biol 1996; 5:1 9. Danielovi V, Adhami J. Mosquitoes of Albania and their medical importance. Ceskoslovenska Parasitologie 1960; 7:41 47 (in Russian). Darsie RF, Jr., Samanidou-Voyadjoglou A. Keys for the identification of the mosquitoes of Greece. J Am Mosq Control Assoc 1997; 13: ECDC. Technical report. Guidelines for the surveillance of invasive mosquitoes in Europe. Stockholm: ECDC, ECDC VBORNET Available at healthtopics/vectors/vectormaps/pages/vbornet_maps.aspx Giatropoulos A. The public health importance of the Asian tiger mosquito. 2013; Giatropoulos A, Emmanouel N, Koliopoulos G, Michaelakis A. A study on distribution and seasonal abundance of Aedes albopictus (Diptera: Culicidae) population in Athens, Greece. J Med Entomol 2012a; 49: Giatropoulos AK, Michaelakis AN, Koliopoulos GTh, Pontikakos CM. Records of Aedes albopictus and Aedes cretinus (Diptera: Culicidae) in Greece from 2009 to Hell Plant Prot J 2012b; 5: Gratz NG. Critical review of the vector status of Aedes albopictus. Med Vet Entomol 2004; 18: Harbach RE. The mosquitoes of the subgenus Culex in southwestern Asia and Egypt (Diptera: Culicidae). Contrib Am Entomol Inst 1998; 24:vi Institut de Veille Sanitaire (INVS). Chikungunya et dengue Données de la surveillance renforcée. France métropolitaine en ; Avaliable at Maladies-infectieuses/Maladies-a-transmission-vectorielle/ Chikungunya/Donnees-epidemiologiques/France-metropolitaine/ Chikungunya-et-dengue-Donnees-de-la-surveillance-renforceeen-France-metropolitaine-en-2014 Lee DJ, Hicks MM, Debenham ML, Griffiths M, et al. The Culicidae of the Australasian Region.Canberra: Australian Government Publishing Service, LifeConops. Presence of Aedes albopictus (Asian tiger mosquito) in Greece. 2015; By conops, on July 21, Available at Lytra ICh, Emmanouel NG. Presence of Culex tritaeniorhynchus(diptera: Culicidae) in rice fields of Western Greece. Hell Plant Prot J 2014; 7: Marchand E, Prat C, Jeannin C, Lafont E, et al. Autochthonous case of dengue in France, October Eurosurveillance 2013; 18:6. Marka A, Diamantidis A, Papa A, Valiakos G, et al. West Nile Virus State of the Art Report of MALWEST Project. Int J Environ Res Public Health 2013; 10:

7 RECORDS OF A. ALBOPICTUS AND Cx. TRITAENIORHYNCHUS EXPANSION IN GREECE 223 Patsoula E, Samanidou-Voyadjoglou A, Spanakos G, Kremastinou J, et al. Molecular and morphological characterization of Aedes albopictus in northwestern Greece and differentiation from Aedes cretinus and Aedes aegypti. J Med Entomol 2006; 43: Patsoula E, Vakali A, Balatsos G, Pervanidou D, et al. West Nile Virus Circulation in Mosquitoes in Greece ( ). Biomed Res Int 2016;2016: Paty MC, Six C, Charlet F, Heuzé G, et al. Large number of imported chikungunya cases in mainland France, 2014: A challenge for surveillance and response. EuroSurveillance 2014; 19: Porter CH, Collins FH. Species-diagnostic differences in a ribosomal DNA internal transcribed spacer from the sibling species Anopheles freeborni and Anopheles hermsi (Diptera: Culicidae). Am J Trop Med Hyg 1991; 45: Samanidou A, Harbach R. Culex tritaeniorhynchus Giles, a newly discovered potential vector of arboviruses in Greece. Eur Mosquito Bull 2003; 16: Samanidou A, Harbach RE. Keys to the adult female mosquitoes (Culicidae) of Greece. Eur Mosquito Bull 2001; 10: Samanidou-Voyadjoglou A, Patsoula E, Spanakos G, Vakalis NC. Confirmation of Aedes albopictus (Skuse) (Diptera: Culicidae) in Greece. Eur Mosquito Bull 2005; 19: Schaffner F, Bouletreau B, Gillet B, Guilloteau G, et al. Aedes albopictus (Skuse, 1894) established in metropolitan France. Eur Mosquito Bull 2001; 9:1 3. Self L, Shin H, Kim K, Lee K, et al. Ecological studies on Culextritaeniorhynchusas a vector of Japanese encephalitis. Bull World Health Organ 1973; 49: Semenza JC, Zeller H. Integrated surveillance for prevention and control of emerging vector-borne diseases in Europe. Euro Surveill 2014; 19:pii: Simon C, Frati F, Beckenbach A, Crespi B, et al. Evolution, weighting, and phylogenetic utility of mitochondrial gene sequences and a compilation of conserved polymerase chain reaction primers. Ann Entomol Soc Am 1994; 87: Toma T, Miyagi I, Crabtree MB, Miller BR. Identification Culex vishnui subgroup (Diptera:Culicidae) Mosquitoes from the Ryukyu Archipelago, Japan: Development of a speciesdiagnostic polymerase chain reaction assay based on sequence variation in ribosomal DNA spacers. J Med Entomol 2000; 37: Walter Reed Biosystematics Unit Smithsonian Institute. Maryland: Silver Spring. AD/CXtri_hab.html 2016 Wesson DM, Porter CH, Collins FH. Sequence and secondary structure comparisons of ITS rdna in mosquitoes (Diptera: Culicidae). Mol Phylogenet Evol 1992; 1: Address correspondence to: Patsoula Eleni Department of Parasitology Entomology and Tropical Diseases National School of Public Health 196, Alexandras Avenue Athens Greece epatsoula@esdy.edu.gr

WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, Oct

WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, Oct HELLENIC CENTRE FOR DISEASE CONTROL AND PREVENTION (HCDCP-KEELPNO) MINISTRY OF HEALTH WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, 2012-19 Oct. 2012 - This weekly epidemiological report

More information

WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, Aug

WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, Aug HELLENIC CENTRE FOR DISEASE CONTROL AND PREVENTION (HCDCP-KEELPNO) MINISTRY OF HEALTH WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, 2012-31 Aug. 2012 - This weekly epidemiological report

More information

ANNUAL EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, 2012

ANNUAL EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, 2012 ANNUAL EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, 2012 This report aims to present an overview of the epidemiological characteristics of the reported cases of West Nile Virus (WNV) infection

More information

WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, Sept

WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, Sept HELLENIC CENTRE FOR DISEASE CONTROL AND PREVENTION (HCDCP-KEELPNO) MINISTRY OF HEALTH WEEKLY EPIDEMIOLOGICAL REPORT WEST NILE VIRUS INFECTION, GREECE, 2012-7 Sept. 2012 - This weekly epidemiological report

More information

Sheep and Goat Pox in Greece - SCoFCAH, Brussels 3-4 July 2014

Sheep and Goat Pox in Greece - SCoFCAH, Brussels 3-4 July 2014 Sheep and Goat Pox in Greece - SCoFCAH, Brussels 3-4 July 2014 HELLENIC REPUBLIC MINISTRY OF RURAL DEVELOPMENT AND FOOD DIRECTORATE GENERAL OF VETERINARY SERVICES ANIMAL HEALTH DIRECTORATE DEPT. OF INFECTIOUS

More information

2. Sampling method There were two types of mosquito trap used in this study. One was the

2. Sampling method There were two types of mosquito trap used in this study. One was the Overseas Practice on(field Epidemiology Collaborative Research) report form(for Student) 2015/05/11 (Year/Month/Day) Name Laboratory Year (Grade) Place of practice Paulina Duhita Anindita Division of Molecular

More information

RESULTS FACT SHEET INDICATOR - SET13: COMPOSITION OF GROSS VALUE ADDED (GVA) PER SECTOR

RESULTS FACT SHEET INDICATOR - SET13: COMPOSITION OF GROSS VALUE ADDED (GVA) PER SECTOR RESULTS FACT SHEET INDICATOR - SET13: COMPOSITION OF GROSS VALUE ADDED (GVA) PER SECTOR DEFINITION- OBJECTIVE This indicator records the composition of the Gross Value Added (GVA) per Prefecture/Region

More information

WEEKLY REPORT. Anopheles fransiscanus Anopheles freebornii Culex erythrothorax

WEEKLY REPORT. Anopheles fransiscanus Anopheles freebornii Culex erythrothorax WEEKLY REPORT OWENS VALLEY MOSQUITO ABATEMENT PROGRAM MAMMOTH LAKES MOSQUITO ABATEMENT DISTRICT WEEK 4: SEPTEMBER 3 TO OCTOBER 6, 218 PAST WEEK S TREATMENT SUMMARY Larvicide treatments are wrapping up

More information

The results of indicators, the reports, and generally the Observatory material are available on the internet at:

The results of indicators, the reports, and generally the Observatory material are available on the internet at: Regional Gross Domestic Product (GDP) 2003 in the Regions of the European Union (EU25) the profile of Regions and Prefectures crossed by the Egnatia motorway The EUROSTAT, the statistical service of the

More information

Weekly Epidemiological Report for West Nile Virus disease, Greece, September

Weekly Epidemiological Report for West Nile Virus disease, Greece, September Weekly Epidemiological Report for West Nile Virus disease, Greece, 2018-27 September 2018 - This weekly epidemiological report aims to present an overview of the reported cases and public health response

More information

Sheep and Goat Pox in Greece (Short overview update as at March 2015)

Sheep and Goat Pox in Greece (Short overview update as at March 2015) HELLENIC REPUBLIC MINISTRY OF PRODUCTION RECONSTRUCTION ENERGY & ENVIRONMENT DIRECTORATE GENERAL OF SUSTAINABLE ANIMAL PRODUCTION & VETERINARY MEDICINE ANIMAL HEALTH DIRECTORATE DEPT. OF INFECTIOUS DISEASES

More information

PRESS RELEASE. Graph 1: Road traffic accidents and casualties, January January Jul-14. Jul-15. Apr-15. Jan-15. Oct-15.

PRESS RELEASE. Graph 1: Road traffic accidents and casualties, January January Jul-14. Jul-15. Apr-15. Jan-15. Oct-15. Jan-11 Apr-11 Jul-11 Oct-11 Jan-12 Apr-12 Jul-12 Oct-12 Jan-13 Apr-13 Jul-13 Oct-13 Jan-14 Apr-14 Jul-14 Oct-14 Jan-15 Apr-15 Jul-15 Oct-15 Jan-16 Apr-16 Jul-16 Oct-16 Jan-17 Apr-17 Jul-17 Oct-17 Jan-18

More information

FAO-EuFMD/EC/OIE Tripartite Group Meeting on control of FMD and other exotic diseases in the Southern Balkans

FAO-EuFMD/EC/OIE Tripartite Group Meeting on control of FMD and other exotic diseases in the Southern Balkans FAO-EuFMD/EC/OIE Tripartite Group Meeting on control of FMD and other exotic diseases in the Southern Balkans GREECE Chryssoula Dile DVM,Msc ANIMAL HEALTH DIRECTORATE DIRECTORATE GENERAL OF SUSTAINABLE

More information

WORKING PAPER In the context of the placement in the Observatory of the Egnatia Odos A.E.

WORKING PAPER In the context of the placement in the Observatory of the Egnatia Odos A.E. COMPARATIVE ANALYSIS OF POPULATION CHANGES IN THE AREAS CROSSED BY THE THREE BASIC ROAD AXES OF GREECE: THE EGNATIA MOTORWAY, THE PA.TH.E. AND THE IONIAN MOTORWAY WORKING PAPER In the context of the placement

More information

Bluetongue emergence in Peloponnisos (Greece) - SCoFCAH, Brussels 3-4 July 2014

Bluetongue emergence in Peloponnisos (Greece) - SCoFCAH, Brussels 3-4 July 2014 HELLENIC REPUBLIC MINISTRY OF RURAL DEVELOPMENT AND FOOD DIRECTORATE GENERAL OF VETERINARY SERVICES ANIMAL HEALTH DIRECTORATE Bluetongue Emergence in Peloponnisos (Greece) (June early July 2014) Dimitrios

More information

MACEDONIA IN GREEK ADMINISTRATION

MACEDONIA IN GREEK ADMINISTRATION 24 M A C E D O N I A - ª π - M A K E D O N I J A MACEDONIA IN GREEK ADMINISTRATION The Balkan Wars of 1912-13 freed Macedonia from Ottoman suzerainty. Its greater part over 50 per cent of the former Ottoman

More information

PRESS RELEASE. ARRIVALS OF NON-RESIDENTS IN GREECE: January - June 2016 HELLENIC REPUBLIC HELLENIC STATISTICAL AUTHORITY. Piraeus, 13 October 2016

PRESS RELEASE. ARRIVALS OF NON-RESIDENTS IN GREECE: January - June 2016 HELLENIC REPUBLIC HELLENIC STATISTICAL AUTHORITY. Piraeus, 13 October 2016 HELLENIC REPUBLIC HELLENIC STATISTICAL AUTHORITY Piraeus, 13 October 2016 PRESS RELEASE ARRIVALS OF NON-RESIDENTS IN GREECE: January - June 2016 According to the Frontier Statistical Survey conducted by

More information

Seasonal prevalence and container preferences of Aedes albopictus in Santo Domingo City, Dominican Republic

Seasonal prevalence and container preferences of Aedes albopictus in Santo Domingo City, Dominican Republic 208 Journal of Vector Ecology December, 2003 Seasonal prevalence and container preferences of Aedes albopictus in Santo Domingo City, Dominican Republic Carlos J. Pena 1, Guillermo Gonzalvez 1, and Dave

More information

Emerging arboviral diseases in Europe The French experience

Emerging arboviral diseases in Europe The French experience Emerging arboviral diseases in Europe The French experience Dr Marie-Claire Paty Coordinator vector born diseases surveillance Santé publique France Annual meeting for infectious disease physicians and

More information

monitoring of egnatia motorways spatial impacts

monitoring of egnatia motorways spatial impacts monitoring of egnatia motorways spatial impacts MINISTRY OF ENVIRONMENT, PHYSICAL PLANNING AND PUBLIC WORKS 1 A TOOL AT THE SERVICE OF SPATIAL DEVELOPMENT The Egnatia motorway Observatory monitors, and

More information

SHIP SANITATION TRAINING NETWORK No

SHIP SANITATION TRAINING NETWORK No SHIP SANITATION TRAINING NETWORK No 2007206 Christos Hadjichristodoulou SHIPSAN TRAINET Project leader Workshop on Generic Preparedness (GP) projects Luxemburg, 30 September 1 October 2009 European Commission

More information

Mosquitoes of the Tokara Archipelago, Japan -I. , Hiroshi Suzuki and Takao Okazawa 4

Mosquitoes of the Tokara Archipelago, Japan -I. , Hiroshi Suzuki and Takao Okazawa 4 18 Mosquito Systematics Vol. 15(l) 1983 Mosquitoes of the Tokara Archipelago, Japan -I.2 Ichiro Miyagi, Takako Toma 2.3, Hiroshi Suzuki and Takao Okazawa 4 ABSTRACT: Nineteen mosquito species belonging

More information

What is VectorNet? Network project + More than just the extension of VBORNET. 21 partners from 14 countries Time-frame:

What is VectorNet? Network project + More than just the extension of VBORNET. 21 partners from 14 countries Time-frame: VectorNet Pirbright Institute Bulent Alten European Network for sharing data on geographic distribution of arthropod vectors transmitting human and animal disease agents What is VectorNet? Network project

More information

Project Concept Note

Project Concept Note North-East Asian Subregional Programme for Environmental Cooperation (NEASPEC) 1. Overview 1. Project Title 2. Goals Project Concept Note Study on Transborder Movement of Amur Tigers and Leopards using

More information

RESULTS FACT SHEET SE-S-2: ACCESSIBILITY OF INDUSTRIAL AREAS

RESULTS FACT SHEET SE-S-2: ACCESSIBILITY OF INDUSTRIAL AREAS RESULTS FACT SHEET SE-S-2: ACCESSIBILITY OF INDUSTRIAL AREAS DEFINITION - OBJECTIVE The indicator records data relevant to Industrial Areas (VIPE) and also other types of production and development related

More information

ABSENCE OF CIRCULATION OF BLUETONGUE VIRUS SEROTYPE 8 IN CAMPO DE GIBRALTAR LVU SINCE NOVEMBER 2010

ABSENCE OF CIRCULATION OF BLUETONGUE VIRUS SEROTYPE 8 IN CAMPO DE GIBRALTAR LVU SINCE NOVEMBER 2010 ABSENCE OF CIRCULATION OF BLUETONGUE VIRUS SEROTYPE 8 IN CAMPO DE GIBRALTAR LVU SINCE NOVEMBER 21 SCoFCAH 15th January 213 BTV-8 8 EPIDEMIOLOGICAL EVOLUTION IN SPAIN First incursion: January 28 Origin:

More information

SPECIES COMPOSITION AND SEASONAL DYNAMICS OF MOSQUITOES (DIPTERA: CULICIDAE) IN FLOODED AREAS OF VOJVODINA, SERBIA

SPECIES COMPOSITION AND SEASONAL DYNAMICS OF MOSQUITOES (DIPTERA: CULICIDAE) IN FLOODED AREAS OF VOJVODINA, SERBIA Arch. Biol. Sci., Belgrade, 62 (4), 1191-1206, 2010 DOI:10.2298/ABS1004206V SPECIES COMPOSITION AND SEASONAL DYNAMICS OF MOSQUITOES (DIPTERA: CULICIDAE) IN FLOODED AREAS OF VOJVODINA, SERBIA A. VUJIĆ,

More information

New Zealand, together with Hawaii,

New Zealand, together with Hawaii, Environmental Health Exotic mosquitoes in New Zealand: a review of species intercepted, their pathways and ports of entry José G. B. Derraik Ecology and Health Research Centre, Department of Public Health,

More information

Inspection system for in use pesticide application equiupment in Greece. First three years of application

Inspection system for in use pesticide application equiupment in Greece. First three years of application Inspection system for in use pesticide application equiupment in Greece. First three years of application G. Bourodimos 1,2, A. T. Balafoutis 3, M. Giamouri 1, S. Fountas 2 1 Hellenic Agricultural Organisation

More information

ENGAGING ALUMNI WORLDWIDE

ENGAGING ALUMNI WORLDWIDE ENGAGING ALUMNI WORLDWIDE COLUMBIA GLOBAL CENTERS and COLUMBIA ALUMNI ASSOCIATION Columbia University Senate Feb. 27, 2015 Mission Statement: Columbia Global Centers Columbia Global Centers promote and

More information

THE MOSQUITO FAUNA OF ROTA ISLAND, MARIANA ISLANDS (DIPTERA: CULICIDAE)

THE MOSQUITO FAUNA OF ROTA ISLAND, MARIANA ISLANDS (DIPTERA: CULICIDAE) J. Med. Entomol. ol. 14, no. 4:411-416 :24 December 1977 THE MOSQUITO FAUNA OF ROTA ISLAND, MARIANA ISLANDS (DIPTERA: CULICIDAE) Wesley R. Nowell and Donald R. Sutton 2 Abstract: A mosquito survey on Rota

More information

Preventing disease Promoting and protecting health

Preventing disease Promoting and protecting health Preventing disease Promoting and protecting health CARPHA A New Caribbean Public Health Agency: Responding to public health emergencies Dr Babatunde Olowokure Director Surveillance, Disease Prevention

More information

WORLDWIDE WATERPROOFING REFERENCES

WORLDWIDE WATERPROOFING REFERENCES WORLDWIDE WATERPROOFING REFERENCES Liquid Applied Polyurethane Waterproofing Systems Professional waterproofing applicators in over 45 countries use the MARISEAL SYSTEM for demanding waterproofing EUROPE

More information

Survey of Japanese Encephalitis Virus in Pigs on Miyako, Ishigaki, Kume, and Yonaguni Islands in Okinawa, Japan

Survey of Japanese Encephalitis Virus in Pigs on Miyako, Ishigaki, Kume, and Yonaguni Islands in Okinawa, Japan Jpn. J. Infect. Dis., 62, 220-224, 2009 Short Communication Survey of Japanese Encephalitis Virus in Pigs on Miyako, Ishigaki, Kume, and Yonaguni Islands in Okinawa, Japan Minoru Nidaira*, Katsuya Taira,

More information

ABSENCE OF CIRCULATION OF BLUETONGUE VIRUS SEROTYPE 8 IN THE SPANISH MAINLAND SINCE FEBRUARY 2009

ABSENCE OF CIRCULATION OF BLUETONGUE VIRUS SEROTYPE 8 IN THE SPANISH MAINLAND SINCE FEBRUARY 2009 ABSENCE OF CIRCULATION OF BLUETONGUE VIRUS SEROTYPE 8 IN THE SPANISH MAINLAND SINCE FEBRUARY 2009 SCoFCAH 7-88 May 2012 BTV-8 8 EPIDEMIOLOGICAL EVOLUTION IN SPAIN First incursion: January 2008 Origin:

More information

Tourism In the Maldives

Tourism In the Maldives Tourism In the Maldives Content 1. Introduction 2. Maldives as a Tourist Destination (Sustainable Development Policies) 3. Marketing and Promotion 4. Tourism Sector Performance 1. INTRODUCTION The island

More information

More information at

More information at Report Information More information at https://www.htfmarketreport.com/reports/1266534 Global Flight Control Computer Market Research Report 2018 Report Code: HTF1266534 Pages: 91 Price: 1-User PDF : $

More information

Oviposition Activity and Seasonal Pattern of a Population of Aedes (Stegomyia) aegypti (L.) (Diptera: Culicidae) in Subtropical Argentina

Oviposition Activity and Seasonal Pattern of a Population of Aedes (Stegomyia) aegypti (L.) (Diptera: Culicidae) in Subtropical Argentina Mem Inst Oswaldo Cruz, Rio de Janeiro, Vol. 98(): 69-663, July 23 69 Oviposition Activity and Seasonal Pattern of a Population of Aedes (Stegomyia) aegypti (L.) (Diptera: Culicidae) in Subtropical Argentina

More information

Rapid detection and evolutionary analysis of Legionella pneumophila serogroup 1 ST47

Rapid detection and evolutionary analysis of Legionella pneumophila serogroup 1 ST47 Rapid detection and evolutionary analysis of Legionella pneumophila serogroup 1 ST47 M. Mentasti, P. Cassier, S. David, C. Ginevra, L. Gomez-Valero, A. Underwood, B. Afshar, J. Etienne, Julian Parkhill,

More information

Sizing up Australia s eastern Grey Nurse Shark population

Sizing up Australia s eastern Grey Nurse Shark population Image: David Harasti A new estimate of adult population size for Australia s eastern Grey Nurse Shark drew on widespread genetic sampling and forensic exploration of family trees. Grey Nurse Sharks are

More information

WORLDWIDE WATERPROOFING REFERENCES

WORLDWIDE WATERPROOFING REFERENCES WORLDWIDE WATERPROOFING REFERENCES Liquid Applied Polyurethane Waterproofing Systems Professional waterproofing applicators in over 70 countries use the MARISEAL SYSTEM for demanding waterproofing EUROPE

More information

WORLD PRESS FREEDOM INDEX 2012

WORLD PRESS FREEDOM INDEX 2012 WORLD PRESS FREEDOM INDEX 2012 1 Finland -10,00 = - Norway -10,00 = 3 Estonia -9,00 - Netherlands -9,00 5 Austria -8,00 6 Iceland -7,00 - Luxembourg -7,00 8 Switzerland -6,20 9 Cape Verde -6,00 10 Canada

More information

HELLENIC NAVY HYDROGRAPHIC SERVICE. XV th MEDITERRANEAN AND BLACK SEAS HYDROGRAPHIC COMMISSION CONTRIBUTION OF GREECE

HELLENIC NAVY HYDROGRAPHIC SERVICE. XV th MEDITERRANEAN AND BLACK SEAS HYDROGRAPHIC COMMISSION CONTRIBUTION OF GREECE HELLENIC NAVY HYDROGRAPHIC SERVICE XV th MEDITERRANEAN AND BLACK SEAS HYDROGRAPHIC COMMISSION CONTRIBUTION OF GREECE Malta, 22-24 October 2007 1. Hydrographic Office: Hellenic Navy Hydrographic Service

More information

possession rate of vehicles and basic developmental indicators in the impact zone of Egnatia motorway, Greece THESSALONIKI, Oct.

possession rate of vehicles and basic developmental indicators in the impact zone of Egnatia motorway, Greece THESSALONIKI, Oct. Αύγουστος 27 possessiion rate off vehiiclles and basiic devellopmentall iindiicators iin the iimpact zone off Egnatiia motorway,, Greece THESSALONIKI, Oct. 27 The present report gives an overview of the

More information

Madera County Grand Jury Final Report Madera County Mosquito and Vector Control District

Madera County Grand Jury Final Report Madera County Mosquito and Vector Control District 2010-2011 Madera County Grand Jury Final Report Madera County Mosquito and Vector Control District INTRODUCTION: 2010-2011 Madera County Grand Jury Final Report Madera County Mosquito and Vector Control

More information

2018 POST SHOW REPORT

2018 POST SHOW REPORT 2018 POST SHOW REPORT Supported by Cooperated with Exhibitor Record-Breaking Participation 1.217 companies and company representatives 43 countries Top 10 exhibiting countries among companies and company

More information

ROMANIA. Table 1. FDI flows in the host economy, by geographical origin. (Millions of US dollars)

ROMANIA. Table 1. FDI flows in the host economy, by geographical origin. (Millions of US dollars) Table 1. FDI flows in the host economy, by geographical origin World.... 2 196 6 435 6 485 11 333 9 928 13 910 4 843 2 939 2 522 2 746 Developed economies.... 1 521 5 361 6 309 11 445 9 136 13 422 4 812

More information

Northern Pacific Sea Star

Northern Pacific Sea Star Northern Pacific Sea Star The Northern Pacific Seastar http://www.marine.csiro.au/leafletsfolder/01npseastar.html Reproduced with permission of Craig Macaulay, CSIRO http://www.marine.csiro.au/leafletsfolder/01npseastar.html

More information

SILK ROAD. A view from the world

SILK ROAD. A view from the world 1 SILK ROAD A view from the world RESPONDENT PROFILE International news consumers from BBC s global panels; Travellers from UNWTO s tour operators database 32% 68% 51 years US$2,160 per person GENDER AGE

More information

22 PEW RESEARCH CENTER. Topline Results. Pew Research Center Spring 2014 survey October 16, 2014 Release

22 PEW RESEARCH CENTER. Topline Results. Pew Research Center Spring 2014 survey October 16, 2014 Release 22 Topline Results Pew Research Center Spring 2014 survey October 16, 2014 Release Methodological notes: Survey results are based on national samples. For further details on sample designs, see Survey

More information

Molecular characterization of Italian Soil-borne cereal mosaic virus isolates

Molecular characterization of Italian Soil-borne cereal mosaic virus isolates 11 Parasitica, 2005, 61(1):11-16 Molecular characterization of Italian Soil-borne cereal mosaic virus isolates C. RATTI 1, A. PISI 1, V. VALLEGA 2 & C. RUBIES AUTONELL 1 1 Department of Agroenvironmental

More information

Membership & Voting Strength - 1 October September 2020

Membership & Voting Strength - 1 October September 2020 ship & Voting Strength - 1 October 2016-30 September 2020 Assessment Period - 1 October 2012-30 September 2016 Junior Team Championships Junior - Championships Junior - Championships Junior - Africa 1

More information

RESUME. Designation : Assistant Professor. Office Address : Jamal Mohamed College Trichy 20

RESUME. Designation : Assistant Professor. Office Address : Jamal Mohamed College Trichy 20 RESUME Name : P. RAJASEKAR Designation : Assistant Professor Department : Zoology Date of Birth : 12.05.1984 Office Address : Jamal Mohamed College Trichy 20 Blood Group : O+ Contact : Mobile +91-99438

More information

Mosquito Control Report 2014 On the Ni-les tun unit of the USFWS Bandon Marsh

Mosquito Control Report 2014 On the Ni-les tun unit of the USFWS Bandon Marsh Mosquito Control Report 2014 On the Ni-les tun unit of the USFWS Bandon Marsh Prepared Coos County Public Health Narrative Rick Hallmark and Nikki Harris December 2014 Introduction Page 1 Monitoring Page

More information

Presentation Title: Aerodromes Licensing Requirements

Presentation Title: Aerodromes Licensing Requirements Presentation Title: Aerodromes Licensing Requirements Presenter s name: Ms Mary-Ann Joubert Manager: Aerodrome Operations Date: 29 March 2017 Content Establishment of ICAO Chicago Convention Members Status

More information

Global Travel Trends 2005

Global Travel Trends 2005 Preliminary World Travel Monitor Results from IPK international for the ITB Berlin Message, 03/10/06 Global Travel Trends 2005 Based on the new data from the 2005 World Travel Monitor, and as it does every

More information

A Medical Mystery of Epidemic Proportions

A Medical Mystery of Epidemic Proportions STO-116 A Medical Mystery of Epidemic Proportions Daphne s Blog - Sunday I m not sure my decision to be a Peace Corp volunteer was a good idea. I thought I was prepared for working in a village where extreme

More information

East Africa s Largest Int l. Trade Exhibition & Conference on Power & Energy

East Africa s Largest Int l. Trade Exhibition & Conference on Power & Energy 07 th 2 0 1 8 East Africa s Largest Int l. Trade Exhibition & Conference on Power & Energy POST SHOW REPORT 29-31 May 2018 KICC, Nairobi, Kenya EXPOGROUP Supports The GO GREEN Campaign Across The Globe.

More information

5.3. Cannabis: Wholesale and Street Prices and Purity Levels

5.3. Cannabis: Wholesale and Street Prices and Purity Levels 5. Prices: Cannabis 5.3. Cannabis: Wholesale and Street Prices and Purity Levels HERBAL CANNABIS Africa East Africa Ghana 51.0 39.2-62.7 1999 Kenya 0.2 0.1-0.3-2002 119.8-1996 Madagascar 1.3 1.1-1.4 100.0

More information

MONTHLY NATURAL GAS SURVEY. November 2009

MONTHLY NATURAL GAS SURVEY. November 2009 MONTHLY NATURAL GAS SURVEY November 2009 INTERNATIONAL ENERGY AGENCY INTERNATIONAL ENERGY AGENCY Next Release: 12 March 2010 MONTHLY NATURAL GAS SURVEY - 1 CONTENTS TABLE 1 Natural Gas Balances in OECD

More information

Outlook Prospects of inbound tourism to Greece in March Evangelia Lamprou. Dr. Aris Ikkos, ISHC.

Outlook Prospects of inbound tourism to Greece in March Evangelia Lamprou. Dr. Aris Ikkos, ISHC. Outlook 2018 Prospects of inbound tourism to Greece in 2018 March 2018 Dr. Aris Ikkos, ISHC Serafeim Koutsos Research Director Analyst ΙΝΣΕΤΕ Republishing is permitted provided the source is mentioned

More information

Austrade Supporting Australian Rail Industry s Global Focus. Rail Sector Lead and Trade Manager, Infrastructure

Austrade Supporting Australian Rail Industry s Global Focus. Rail Sector Lead and Trade Manager, Infrastructure Austrade Supporting Australian Rail Industry s Global Focus Peter Ironmonger Rail Sector Lead and Trade Manager, Infrastructure Austrade s network global local knowledge and relationships Helping your

More information

The Geography of the Middle East

The Geography of the Middle East The Geography of the Middle East Unit 1 Focus The focus of this unit is how physical and cultural geography shape the environment, nations, and people of the Middle East The Middle East is also called

More information

Paragonimus mexicanus Miyazaki e Ishii, 1968

Paragonimus mexicanus Miyazaki e Ishii, 1968 Morphological and molecular study of three populations of Miyazaki e Ishii, 1968 (Digenea: Paragonimidae) in Mexico Jorge López Caballero 1, Virginia León Règagnon 1, Luis García Prieto 1, David Osorio

More information

Outlook for air travel markets

Outlook for air travel markets Outlook for air travel markets June 2016 Brian Pearce Chief Economist International Air Transport Association Extended period of strong air travel market growth 30% 25% 20% 15% 10% 5% 0% -5% -10% -15%

More information

Outlook Update

Outlook Update Outlook 2018 - Update Prospects of inbound tourism to Greece in 2018 2 nd edition Apr. 2018 Dr. Aris Ikkos, ISHC Serafeim Koutsos Research Director Analyst ΙΝΣΕΤΕ Republishing is permitted provided the

More information

INTERNATIONAL TRAVEL AND TOURISM

INTERNATIONAL TRAVEL AND TOURISM INTERNATIONAL TRAVEL AND TOURISM YEAR 2014 1. INTRODUCTION This issue of the Economic and Social Indicators presents data on International Travel and Tourism for the year 2014. A brief on the compilation

More information

Accounting Basis by Country

Accounting Basis by Country Accounting Basis by Country Egypt Africa Northern Africa Morocco Africa Northern Africa Sudan Africa Northern Africa Western Sahara Africa Northern Africa Angola Africa Sub-Saharan Africa Benin Africa

More information

The Silk Road: Unlocking the Tourism potential through collaboration

The Silk Road: Unlocking the Tourism potential through collaboration EURAC Research Bozen/Bolzano, 31 May 2018 The Silk Road: Unlocking the Tourism potential through collaboration Alla Peressolova aperessolova@gmail.com why tourism & travel? 1.323 mln+7% travellers 2017

More information

The W20. The 20 wealthiest countries in the world. Publication Date: October 2015 NEW WORLD WEALTH

The W20. The 20 wealthiest countries in the world. Publication Date: October 2015 NEW WORLD WEALTH The W20 The 20 wealthiest countries in the world Publication Date: October 2015 W20 wealth rankings The following tables ranks the top 20 countries in the world by total individual wealth held. Total individual

More information

Financial Reporting Standards Adoption by Country

Financial Reporting Standards Adoption by Country Financial Reporting Standards Adoption by Country Egypt Africa Northern Africa Morocco Africa Northern Africa Sudan Africa Northern Africa Western Sahara Africa Northern Africa Angola Africa Sub-Saharan

More information

CAPSCA Global Feedback

CAPSCA Global Feedback CAPSCA Global Feedback CAPSCA-MID/06 February 2017 Dr. Ansa Jordaan Chief, Aviation Medicine Section International Civil Aviation Organization Public Health Effects on Aviation Spread of disease by air

More information

SECURITY INDUSTRY MET AT ISAF FOR THE 19 th TIME

SECURITY INDUSTRY MET AT ISAF FOR THE 19 th TIME SECURITY INDUSTRY MET AT ISAF FOR THE 19 th TIME ISAF EXHIBITION ORGANIZED ON 17 th -20 th SEPTEMBER 2015 BROUGHT TOGETHER THE SECURITY SECTOR AGAIN International ISAF Exhibitions, the Only Exhibition

More information

INDICATOR RESULTS FACT SHEET SET08: ACCESSIBLE SITES OF CULTURAL AND TOURIST INTEREST

INDICATOR RESULTS FACT SHEET SET08: ACCESSIBLE SITES OF CULTURAL AND TOURIST INTEREST INDICATOR RESULTS FACT SHEET SET08: ACCESSIBLE SITES OF CULTURAL DEFINITION - OBJECTIVE The indicator determines the important areas of tourist interest of Zone ΙV that are accessible via an interchange

More information

North Africa and Southwest Asia. Economic Geography

North Africa and Southwest Asia. Economic Geography North Africa and Southwest Asia Economic Geography Natural Resources Oil is the major resource in this region. Many countries in this region are members of OPEC (Organization of Petroleum Exporting Countries).

More information

INTERNATIONAL TRAVEL AND TOURISM

INTERNATIONAL TRAVEL AND TOURISM INTERNATIONAL TRAVEL AND TOURISM JANUARY TO SEPTEMBER 2017 1. INTRODUCTION This issue of the Economic and Social Indicators presents data on International Travel and Tourism for the first nine months of

More information

Comoros Island. CITIZENSHIP By INVESTMENT PROGRAM.

Comoros Island. CITIZENSHIP By INVESTMENT PROGRAM. Comoros Island CITIZENSHIP By INVESTMENT PROGRAM www.prandcitizenship.com ABOUT COMOROS ISLAND The official name of the country is the Union of the Comoros. It is a sovereign island nation in the Indian

More information

Domestic Tourism. in Asia and the Pacific

Domestic Tourism. in Asia and the Pacific Domestic Tourism in Asia and the Pacific Table of Contents Foreword 9 Acknowledgement 11 Executive Summary 13 Introduction 41 1 Australia 51 1.1 Domestic Tourism in Australia 51 1.1.1 Introduction 51 1.1.2

More information

The stonefly (Plecoptera) fauna of the Carpathian Basin and the Balkans (Dávid Murányi, Tibor Kovács, Kirill Orci)

The stonefly (Plecoptera) fauna of the Carpathian Basin and the Balkans (Dávid Murányi, Tibor Kovács, Kirill Orci) The stonefly (Plecoptera) fauna of the Carpathian Basin and the Balkans (Dávid Murányi, Tibor Kovács, Kirill Orci) The stonefly fauna of the Carpathians, and especially of the Balkans are rich in comparison

More information

KINGDOM OF CAMBODIA NATION RELIGION KING 3

KINGDOM OF CAMBODIA NATION RELIGION KING 3 KINGDOM OF CAMBODIA NATION RELIGION KING 3 TOURISM STATISTICS REPORT October 2015 MINISTRY OF TOURISM Statistics and Tourism Information Department No. A3, Street 169, Sangkat Veal Vong, Khan 7 Makara,

More information

The World Pasta Industry in 2011

The World Pasta Industry in 2011 The World Pasta Industry in 2011 Survey The World Pasta Industry in 2011 25 October 2012 1 Like every year, we have completed our Annual Survey on the World Pasta Industry. We would like to thank all the

More information

COUNTRY & TERRITORY PROFILES A Summary of Oil Spill Response Arrangements & Resources Worldwide

COUNTRY & TERRITORY PROFILES A Summary of Oil Spill Response Arrangements & Resources Worldwide 3. COUNTRY & TERRITORY PROFILES (including date of update) A Albania August 2010 Antigua January 2009 Algeria May 2011 Argentina December 2015 Angola May 2009 Aruba April 2008 Anguilla November 2005 Australia

More information

Performance Derby: MSCI Share Price Indexes

Performance Derby: MSCI Share Price Indexes Performance Derby: MSCI Share Price Indexes January 5, 2018 Dr. Ed Yardeni 516-972-7683 eyardeni@yardeni.com Joe Abbott 732-497-5306 jabbott@yardeni.com Please visit our sites at blog.yardeni.com thinking

More information

Performance Derby: MSCI Regions/Countries Earnings & Revenues Growth 2018E / 2017E / 2016A

Performance Derby: MSCI Regions/Countries Earnings & Revenues Growth 2018E / 2017E / 2016A Performance Derby: MSCI Regions/Countries Earnings & Revenues Growth 2018E / 2017E / 2016A December 6, 2017 Dr. Ed Yardeni 516-972-7683 eyardeni@yardeni.com Joe Abbott 732-497-5306 jabbott@yardeni.com

More information

Exports of Canadian Grain and Wheat Flour

Exports of Canadian Grain and Wheat Flour Canadian Grain Commission Commission canadienne des grain JANUARY CROP YEAR 2011 Exports of Canadian Grain and Wheat Flour date Tonnes (millions) 8.0 7.0 6.0 5.0 4.0 3.0 2.0 1.0 0.0 4.0 3.5 3.0 2.5 2.0

More information

Ancient Greece. Roots of Western Civilization

Ancient Greece. Roots of Western Civilization Ancient Greece Roots of Western Civilization Greece Map Identification Balkan Peninsula: Cities: Troy Mycenae Knossos Ephesus Delphi Athens Sparta Other Geographic features: Mount Olympus Aegean Sea Pelopennesus

More information

IATA ECONOMIC BRIEFING MARCH 2011

IATA ECONOMIC BRIEFING MARCH 2011 IATA ECONOMIC BRIEFING MARCH 2011 WHAT DRIVES THE SIZE OF PREMIUM AIR TRAVEL MARKETS? WHY PREMIUM AIR TRAVEL IS AN IMPORTANT TRAVEL MARKET SEGMENT The premium (first and business class) travel segment

More information

Global travel patterns: an overview

Global travel patterns: an overview Journal of Travel Medicine, 2017, 1 5 doi: 10.1093/jtm/tax007 Perspective Perspective Global travel patterns: an overview Dirk Glaesser*, John Kester, Hanna Paulose, Abbas Alizadeh, and Birka Valentin

More information

Exports of Canadian Grain and Wheat Flour

Exports of Canadian Grain and Wheat Flour Canadian Grain Commission Commission canadienne des grain FEBRUARY CROP YEAR 2012 Exports of Canadian Grain and Wheat Flour date Tonnes (millions) 10.0 9.0 8.0 7.0 6.0 5.0 4.0 3.0 2.0 1.0 0.0 6.0 5.5 5.0

More information

Exports of Canadian Grain and Wheat Flour

Exports of Canadian Grain and Wheat Flour Canadian Grain Commission Commission canadienne des grain JULY CROP YEAR 2012 Exports of Canadian Grain and Wheat Flour date Tonnes (millions) 14.0 12.0 10.0 8.0 6.0 4.0 2.0 0.0 9.0 8.0 7.0 6.0 5.0 4.0

More information

RESULTS FACT SHEET SE-F-3: ACCESSIBILITY OF SITES OF CULTURAL & TOURIST INTEREST

RESULTS FACT SHEET SE-F-3: ACCESSIBILITY OF SITES OF CULTURAL & TOURIST INTEREST DEFINITION - OBJECTIVE Currently, the indicator records the areas of tourist interest in Zone II (prefectures crossed by the Egnatia motorway) that are accessible via an interchange of the axis and their

More information

Silk Road for Japanese Market. presented by Saiyu Travel Co., Ltd.

Silk Road for Japanese Market. presented by Saiyu Travel Co., Ltd. Silk Road for Japanese Market presented by Saiyu Travel Co., Ltd. About Saiyu Travel Since 1973, We are sending Japanese tourist to a destination of Silk Road, mainly to Xinjiang-China, Central Asia countries,

More information

REPUBLIC OF KOREA. Table 1. FDI flows in the host economy, by geographical origin. (Millions of US dollars)

REPUBLIC OF KOREA. Table 1. FDI flows in the host economy, by geographical origin. (Millions of US dollars) Table 1. FDI flows in the host economy, by geographical origin World 4 130 3 395 4 383 8 980 7 050 4 950 2 629 8 409 7 501 8 117 10 247 11 117 Developed economies 3 123 2 385 2 949 7 185 5 743 3 423-275

More information

Notes on the Mosquito Fauna of the Canary Islands. (Diptera: Culicidae)

Notes on the Mosquito Fauna of the Canary Islands. (Diptera: Culicidae) Mosquito Systematics Vol. W(3) 1980 349 Notes on the Mosquito Fauna of the Canary Islands (Diptera: Culicidae) M. Baez and J. M. Fernandez 1 Department of Zoology University of La Laguna Tenerife, Canary

More information

PATA Strategic Intelligence Centre. Tsunami Recovery Update. Wednesday June 22, 2005

PATA Strategic Intelligence Centre. Tsunami Recovery Update. Wednesday June 22, 2005 PATA Strategic Intelligence Centre Tsunami Recovery Update Wednesday June 22, 2005 Page 1 of 6 Y-T-D Inbound (Source: Relevant NTOs, Airport Authorities) Year Destination Period 2004 2005 Change The Americas

More information

Creative Industries in Greece

Creative Industries in Greece Creative Industries in Greece Alina Hyz Kostas Karamanis Creative Industries in Greece An Empirical Analysis from the Region of Epirus Alina Hyz Piraeus University of Applied Sciences, Greece Kostas Karamanis

More information

What is meant by site?

What is meant by site? Site and Situation What is meant by site? Site is the actual location of a city. Example NYC is located on a large, deep water harbor, next to the Atlantic Ocean. What is meant by situation? Situation

More information

What we have in 2009?

What we have in 2009? Academic Ranking of World Universities 2009 Dr. Ying CHENG & Prof. Nian Cai LIU Center for World Class Universities Graduate Schoolofof Education Shanghai Jiao Tong University Third International Conference

More information

HELLENIC NAVY HYDROGRAPHIC SERVICE. XVI th MEDITERRANEAN AND BLACK SEAS HYDROGRAPHIC COMMISSION NATIONAL REPORT OF GREECE

HELLENIC NAVY HYDROGRAPHIC SERVICE. XVI th MEDITERRANEAN AND BLACK SEAS HYDROGRAPHIC COMMISSION NATIONAL REPORT OF GREECE HELLENIC NAVY HYDROGRAPHIC SERVICE XVI th MEDITERRANEAN AND BLACK SEAS HYDROGRAPHIC COMMISSION NATIONAL REPORT OF GREECE Odessa - Ukraine, 22-24 September 2009 1. Hydrographic Service: General Hellenic

More information

Highlights of The Travel & Tourism Competitiveness Report 2008

Highlights of The Travel & Tourism Competitiveness Report 2008 Highlights of The Travel & Tourism Competitiveness Report 2008 Switzerland is number one in the Travel & Tourism Competitiveness Index (TTCI) 2008, followed by Austria and Germany, the same as in the TTCI

More information